• Journal of Korean Society of Forest Science
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• v.85 no.3
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• pp.524-531
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• 1996

The objective of this study was to determine the important factors affecting the early natural regeneration of Pinus densiflora S. et Z. Seed germination, survival and height growth of the pine seedlings were examined at four experimental plots of P. densiflora forests including forest roadside. Plots I and III were thinned in 1992, plot II was treated with hexazinone in 1993 after seed tree method was applied in 1982 and 1989. Seedlings emerged after thinning and the rate of their emergence depended on forest floor conditions. Seedling survival was related with light conditions, herbaceous vegetation, and degrees of rainfall. More seedlings emerged in forest hauling roads than within the forest because soil scarification due to logging provided better conditions for seed germination and seedling growth. Seed supply was enough, but germination rate was very low in the forest compared with that in the greenhouse because micro-environments such as organic layer and herbaceous plant acted as limiting factors. Weed control was needed for 4 to 5 years until seedlings grew upto about 50cm in height which was similar to that of herbaceous plants. But 10-40% coverage of herbaceous vegetation was needed to protect small seedlings from heavy rain. Forest works such as thinning and logging, over 40% of light were important factors to help the pine regeneration. However, over 70% coverage of herbaceous vegetation, and heavy rain inhibited rather survival and growth of pine seedlings.

• Korean Journal of Environment and Ecology
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• v.29 no.5
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• pp.785-790
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• 2015

As an alternative to large-scale clear cutting silviculture, strip clear cutting (SC) is being considered as a system compatible with ecological conservation and forest regeneration. In South Korea, application and effectiveness of SCC in varying forest types were rarely found. In this study, under the subject of strip clear cutting lands of pinus densiflora forest at Samcheok, Gangwon-do Province, the developmental aspect of low vegetation prior to and after deforestation and the correlation between environmental factor and pine regeneration were analyzed. The cover rate of understory vegetation was appeared to be increased after deforestation and rapidly increased two years after deforestation, and it was evaluated to be affected by vigorous tree species and photophilic species. From the perspective of relative importance value, Quercus mongolica, Artemisia keiskeana, and Rubus crataegifolius that influence the cover rate showed the inclination of continuous growth. The diversity of species showed increment inclination as well due to introduction and settlement of early transient species. As a result of analyzing the correlation between vegetation and environmental factor and generation of pine tree size, the soil exposure rate, intensity of light, and canopy openness showed positive relationship, and the understory vegetation cover and woody debris cover rate showed negative relationship.

• Journal of Korean Society of Forest Science
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• v.112 no.3
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• pp.331-339
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• 2023

This study was conducted to identify seed inflow characteristics according to harvest typeand to provide basic data for developing a regeneration technology for secondary growth forests in Korean red pine (Pinus densiflora) succession forest formation by natural regeneration. Experimental sites were established by applying seed tree (single and group) and clear-cutting methods (10- 20- 30-m strip and 20- 30- 40-m patch). The seed inflow characteristics of the natural regeneration site were analyzed for 6 years from 2014. Most seeds were flowed to the regeneration stand from October to November. In years with good seed fructification, more than 80% were flowed in October. The average annual seed inflow by harvest type was highest in the seed tree area (296,000 seeds/ha/yr), followed by the 20-m patch clear-cutting area (291,000 seeds/ha/yr) and 10-m strip clear-cutting area (281,000 seeds/ha/yr). The distribution uniformity of seed inflow according to treatment was analyzed in the order of the 20-m strip clear-cutting area (52.2), 20-m patch clear-cutting area (52.9), and 10-m strip clear-cutting area (56.1). As a result, the 10-m strip and 20-m patch clear-cutting areas with relatively small harvest areas showed high seed inflow and distribution uniformity.

• Journal of Korean Society of Forest Science
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• v.96 no.4
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• pp.419-424
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• 2007

The urban forest offer many opportunities to maintain and enhance local and regional biological diversity. Pinus densiflora stands in isolated urban forest in five areas of Seoul, Incheon, Daegu, Ulsan, and Cheongju were investigated by landscape and community ecological methods for diagnosis of current status and construction of conservation plan. Species composition of five study areas revealed regional characteristics. As the result of stand dynamics, absence of pine seedlings and saplings were observed. Cheongju, significantly, revealed highest species diversity (H'=1.86) and richness (24) among study areas, and the opposite was true for Daegu (1.35 for H', and 14 for richness). Ratio of alien species was highest in Daegu (15.8%) (F=5.05, p=0.0016). Based on landscape analysis, complexity of pine patch was higher in local city than metropolitans, but statistically not significant (p=0.133). Complexity of pine patch revealed significant positive correlation with diversity and richness, but not with percentage of aliens. Synthetically, ecological management based on community and landscape ecological information was recommended. Cautious artificial regeneration strategy was necessary to regenerate pine forest in urban area.

• Journal of Korean Society of Forest Science
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• v.78 no.4
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• pp.401-411
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• 1989

The embryos of Pinus taeda, P. rigida, and P. taeda ${\times}$ rigida were cultured for adventitious shoot regeneration in vitro. Culture media were modified from Gresshoff and Doy (MGD), Murashige and Skoog (MMS), Lloyd and McCown (MLM), and Schenk and Hildebrandt (MSH). NAA was added to initiation media at a concentration of 0.1 or 0.01 mg/l. BAP was used at the concentrations of 0.1. 0.5, 1, 2, or 5mg/l. Each explant was induced for 3-4 weeks on solid medium. All explants were cultured up to 16 weeks. Illumination was about $1506{\pm}540lux$ at the level of the tissues in the growth room with a temperature of $25{\pm}2^{\circ}C$. A 16-hour photoperiod per 24 hours was used. Half-strength medium was used for all the subcultures. For shoot production by loblolly pine, MMS, MLM, or MSH is preferred with 5 mg/l BAP with either 0.1 or 0.01 mg/l NAA. For shoot production by pitch pine, MMS, MLM, or MSH is recommended with 2 or 5 mg/l BAP with 0.1 mg/l NAA. For shoot production by the hybrid pine, MMS or MLM is more effective with 1, 2 or 5 mg/l BAP with 0.1 mg/l NAA. There were no differences recognized among the species tried in the patterns of bud formation and shoot development. Different composition of media, in major and minor salts or possibly in vitamins, should be tested for the two developmental stages of adventitious shoots ; the induction of shoot buds and the elongation of them into shoots.

• Korean Journal of Environment and Ecology
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• v.23 no.1
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• pp.56-65
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• 2009

In Korea, Red pine(Pinus densiflora) stands at a very important place, historically, culturally, and emotionally and it is one of the tree species that can produce domestic timber as forest resources economically. The growing space for Red pine forest in Korea is gradually reducing while the space for deciduous tree forests including oak series is gradually increasing. Thus, it is required that the research work should be done on renewal for pine forest retention and its related forest management practices. This research aims at developing stable, sustainable management forests and inducing nature renewal by grasping growth environment and succession process through the pattern of stand and structure analysis of the red pine(Pinus densiflora) for central region. The pine forests in central region were classified into four communities, such as Acer pseudo-sibolianum, Quercus acutissima, Lindera erythrocarpa, and Pinus densiflora and they are showing different characteristics by pattern according to their growing district. There appeared a lot of red pines in the upper layer, but there existed high weight of broad-leaved forest tree species, such as oak series in the middle layer and the appearance of the red pine was meager in the lower layer. Therefore, it appears that the red pine has a high influential ecological strength in terms of correlativity; however, if the red pine in tree layer is dead by aging in nature state with the lapse of time, it is considered that oak species, such as Quercus mongolica, Quercus variabilis, and Quercus serrata will be dominant below sub-tree layer. In addition, there appear differences in characteristics and growth pattern of the red pine forest depending on stand pattern, so it is considered that a differential application method should be employed in the pine forest management.

• Journal of the Korean Society of Environmental Restoration Technology
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• v.2 no.4
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• pp.1-8
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• 1999

Forest devastation in Korea was caused by several factors such as internal factors from geological features and external factors from artificial forest damages including fuel wood collection from forests, forest fires, shifting cultivations and so on. According to the reports of 1935, lots of forest devastation in Kyungpook province area occurred around the main and branch stream of Nakdong river. Main factors of occurring forest devastation in 1935 were investigated by the methods of forest devastation rate and the population density at the basin of Nakdong river. But based on our study, forest devastation mainly occurred in rock zones of granite and granite gneiss, next to Nakdong formation but scarcely occurred in Hayang formation. Clay of the weathered soils of granite and granite gneiss was lost by rainfall, but remaining coarse-sandy soils(or grits) have poored conditions in vegetation's growth, which are due to high level of water permeability, lack of water-holding capacity and dried conditions. Generally, pine forests are mainly growing up in these regions. It is supposed that forest devastation was accelerated due to long periods of natural regeneration and no ability of natural regeneration by sprout after frequent collections of fuel wood and cuttings from pine forest on those grit areas. These results indicated that the high rate of forest devastation occurred around the basin with the high resident population density, which was partly due to forest damages by fuel collection. Moreover, both geological features and number of residents had much influence on forest devastation. Forest devastation was positively correlated with those variables(r=+0.73).

• Plant Biotechnology Reports
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• v.5 no.2
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• pp.177-186
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• 2011

This study describes for the first time in Pinus genus a plant regeneration system via a combined pathway of somatic embryogenesis and organogenesis from immature seeds of radiata pine. Somatic embryos were obtained from embryogenic line 2162 of Pinus radiata D. Don on EDM basal medium containing $60{\mu}M$ ABA and 6% sucrose. The explants used for organogenesis experiments were either freshly collected somatic embryos or somatic embryos germinated for 1 week. Germination medium was half-strength LP medium, supplemented with 0.2% activated charcoal. Different induction periods and BA concentrations were assayed for shoot induction. After induction treatments, explants were elongated on the same medium used for germination stage. Rooting medium was quarter-strength LP medium supplemented with three different auxin treatments: $1.5mg\;L^{-1}$ 1-naphthalene acetic acid (NAA), $1.5mg\;L^{-1}$ indole-3-butyric acid (IBA) and $1mg\;L^{-1}$ IBA with $0.5mg\;L^{-1}$ NAA (MIX). The effect of the photon flux ($120mmol\;m^{-2}\;s^{-1}$ and darkness) in the first week of the explants in the rooting media was also tested. This methodology could offer an alternative to overcome some problems associated with somatic embryogenesis such as the seasonality of embryogenic tissue (ET) initiation or a low embryo production from the ET, a particularly important issue in the case of genetically transformed ETs.

• Journal of Korean Society of Forest Science
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• v.89 no.2
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• pp.185-197
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• 2000

This study was carried out to research structure of forest vegetation and site condition, and supply basic data for conservation of pine forest and ecologically sustainable forest management and control in Anmyondo area. The forest of Anmyondo were largely classified as four forest community : Pinus densiflora community, Pinus thunbergii community, Pinus rigida community, and Quercus variabilis community. Organic matter, total nitrogen, and other nutrients in soil of the study area were relatively low, and average soil pH was 4.9. Soil texture was sandy clay or heavy clay and so the aeration and permeability could be poor. In this study site, the methods such as the promotion of germination of pine seeds through removal of forest floor and inducement of natural regeneration through such method as regeneration under shelter wood would be better than the method such as afforestation after clear-cutting for conservation the pine forests. Also, it will be a good method to practice tending for hardwood species in the dominant areas by hardwood species.

• Journal of Korean Society of Forest Science
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• v.96 no.1
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• pp.107-114
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• 2007

Establishments of the seedlings inside the natural forest from adjacent artificial forests would be an important factor in forest stand dynamics. This study was conducted to see the invasion of Korean pine (Pinus koraiensis) seedlings which is not native in this region, into the natural deciduous broad-leaved forest in Gwangneung, Korea. There is no mother tree at the I ha study site while the number of naturally regenerated P. koraiensis seedlings was 345 trees and 56% of them were clumped with more than two seedlings at each point. Applying the image segmentation method to IKONOS satellite image of January, 2003, the distance from the center of 1 ha study site to the nearest mother tree and plantation of Korean pine were 200 m and 270 m, respectively. The average height and root-collar diameter of the seedlings were 34 em and 7 mm, respectively and the age of 207 seedlings (60%) were below 5 years old. Most abundant range of soil moisture gradient and LAl (leaf area index) were from 16 to 20% and those of LAI were from 3.1 to 3.5. To understand the dynamics and seed dispersal pattern of Korean pine in the Gwangneung natural deciduous broad-leaved forests, additional studies not only long-term monitoring of growth and mortality of naturally regenerated Korean pine seedlings but also application of stable isotope analysis and molecular genetic techniques was recommended.