• Title/Summary/Keyword: Climax stage

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The Development of Climax Index by Analysis of Eco-morphological Characters for Major Deciduous Tree Species

  • Kim, Ji Hong;Chung, Sang Hoon;Lee, Jeong Min;Kim, Se Mi
    • Journal of Forest and Environmental Science
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    • 제28권4호
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    • pp.199-204
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    • 2012
  • This study was conducted to estimate climax index by eco-morphology for major 36 tree and sub-tree species in natural deciduous forests so as to interpret seral position of each species in the forest community. Fourteen eco-morphological characters which were considered to be associated with successional gradient in the forest were selected for the study. Four levels per character for each species were given on a standardized scale of increasing climax, and the index was computed by the proportion of the sum of total scores, expressed by percent values. With calculated mean value of 54.8 for all indices, Carpinus cordata had the highest index value of 90.5, and Populus davidiana recorded the lowest of 13.2. The most climax group, greater than 70 of the index, contained only 8 species, intermediate group, between 41 to 70 of the index, had 23 species, and the most pioneer group, less than 40 of the index comprised 5 species. The result has noticed that the large number of species would take advantage of most diverse resource and niche in the intermediate stage of the sere in the forest. By cluster analysis all 36 species were subjected to be classified into several species groups which had common similar eco-morphological characteristics. The indices were additionally plotted on the two dimensional graph to recognize the positions related to the light absorption factor and reproduction factor. The climax index of tree and sub-tree species developed by this study could be applied to understand the present status of successional stage on the basis of species composition by the method of summing up the indices. And comparison of forest successional stage among various forest communities could be done by summing up the climax indices of composed species in each community. However, this kind of applied methodology should be limited to the forest of similar species composition and site condition.

난온대 상록활엽수림 지역의 식생천이계열 고찰 -육박나무군락을 중심으로- (A Review of Vegetation Succession in Warm-Temperate Evergreen Broad-Leaved Forests -Focusing on Actinodaphne lancifolia Community-)

  • 박석곤;최송현;이상철
    • 한국환경생태학회지
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    • 제32권1호
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    • pp.77-96
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    • 2018
  • 난온대 상록활엽수림의 식생천이계열을 논의하기 위해 극상림이라 알려진 육박나무군락이 생육하는 한국 3곳(비진도, 애도, 보길도)과 일본 1곳(다치바나산)을 조사 및 분석했다. 이를 토대로 한국, 일본, 중국, 대만의 관련문헌을 검토해 상록활엽수림의 분포특성, 식생천이계열 및 극상수종을 고찰했다. 다치바나산과 애도는 육박나무군락이 가장 발달한 식생구조를 보였지만, 토양분석 및 CCA분석에서 육박나무군락을 난온대 천이극상단계의 조건으로 판단하기엔 부족함이 보였다. 한국과 일본에서 육박나무군락은 드물게 분포하는 반면, 동아시아 난온대 지역에 보편적인 식생유형은 후박나무류, 잣밤나무류, 가시나무류였다. 한국 난온대 지역의 식생천이계열은 곰솔 소나무 졸참나무 등(초기단계)${\rightarrow}$후박나무 생달나무 참식나무 육박나무 등(중간단계)${\rightarrow}$구실잣밤나무 붉가시나무 참가시나무 등(극상단계)의 순으로 2차천이가 진행될 것으로 추정된다. 단, 조풍의 영향을 강하게 받는 입지조건이나 잣밤나무류 가시나무류의 종자공급이 어려운 곳에서는 후박나무류 유형이 토지극상일 것이다.

Trichophyton verrucosum var album 의 실험적(實驗的) 감염(感染) 기니픽에 대한 병리학적(病理學的) 관찰(觀察) (Pathological observations of guinea pigs experimentally infected with Trichophyton verrucosum var album)

  • 원종한;조성환
    • 대한수의학회지
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    • 제33권2호
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    • pp.309-320
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    • 1993
  • Guinea pigs were cutaneously inoculated with Trichophyton verrucosum var album, that is a common causative fungus of dermatophytosis in cattle. The developmental process of lesions, clinical and histopathological changes and reisolations of the fungi were studied to evaluate the pathogenicity of Trichophyton verrucosum var album in guinea pigs. Results obtained through the experiments were summarized as follows : 1. The incidence of infection of the clipping group was 13(86%) of 15 animals, and that of the plucking group was 14(93%) of 15 animals. In both of the clipping and plucking groups, visible cutaneoas lesions were developed between 4 and 7 days post inoculation(p.i.). The spreading and the climax stages persisted for 4 to 11 and 6 to 12 days, respectively. 2. In macroscopic observations, formation of various degree of erythemas and scales over the inoculated skin sites were observed in the spreading stage. In the climax stage, exudative changes and dark red crusts were formed as typical circular lesions. In the healing stage, the lesions revealed shedding of crust, alopecia and hair regrowth. 3. In histopathological observations, infiltration of inflammatory cells, hyperplasia, microabscesses and keratinous-hyaloid materials of epidermis were observed in the spreading stage. Hyphal invasion was primarily observed at the level of epidermis and pilosebaceous ducts. In the climax stage, the infected epidermis was thick with severe hyperplasia, hyperkeratosis and acanthosis. The microabscesses with fungal hyphae, folliculitis and hyperplasia of external root sheath were observed in the dermis. The fungal hyphae were observed only in the tissues of hair follicles, that were internal root sheath, cuticle, the keratinized portions of cortex and medulla 4. In reisolation of the inoculated fungus, all trials for ten animals showed positive cultures until 25 days p.i.. Afterward, the reisolation rates were gradually decreased, showing all negative after 40 days p.i..

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북한산국립공원의 현존식생과 잠재자연식생 (Actual Vegetation Potential Natural Vegetation of Pukhansan National Park, Mid-western Korea)

  • 이은복
    • The Korean Journal of Ecology
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    • 제20권6호
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    • pp.439-450
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    • 1997
  • The potential natural vegtation of the Pukhansan National Park area, mid-western Korea, was inferred from the actual vegetation. With the phytosociological classification and field surveys, the actual vegetation map of the park area was made on a scale of 1:25, 000, including fourteen communities. By the analyses of the species diversity, the age structure, the human interferences and various informations on vegetation changes, two pathways of late stage succession from Pinus densiflora forests to the climatic climax were suggested. One is from Quercus serrata forests to Q. mongolica forests throughout the mountain and the other, from Q. variabilis or Q. acutissima forests to Carpinus laxiflora forests in lower parts. Considering the vegetation changes, the potential natural vegetation of the park area mainly composed of Q. mongolica, C. laxiflora, P. densiflora and Zelkova serrata forest as the climatic and/or edaphic climax was inferred.

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난온대 상록수림지역의 식생구조와 천이계열

  • 오구균;최송현
    • The Korean Journal of Ecology
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    • 제16권4호
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    • pp.459-476
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    • 1993
  • 내륙지역(두륜산, 불갑산)과 서남해 섬지역(홍도, 진도)의 상록활엽수림과 주변 낙엽활엽수림을 대상으로 식생구조와 식생천이 계열을 분석, 추정하였다. 4개 지역의 상록활엽수림은 대부분 50여 년생 미만의 영급구조로서 처닝도중상의 군집구조를 나타내고 있었으며, 교목층에서의 우점종은 구실잣밤나무, 후박나무, 붉가시나무, 참식나무, 종가시나무, 참가시나무 등이었다. 상록활엽수림대의 식생은 낙엽활엽수종에서 상록활엽수종으로 식생천이가 진행중이었으며, 교목층에서는 후박나무, 참식나무 및 황칠나무가, 아교목층 및 관목층에서는 식나무와 센달나무가 극상단계의 수종으로 추정되었다.

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오대산 국립공원 노인봉지역 식물군집구조분석 (Plant Community Structure Analysis in Noinbong area of Odaesan National Park)

  • 최송현;권전오;민성환
    • 한국환경생태학회지
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    • 제9권2호
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    • pp.156-165
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    • 1996
  • 극상림의 군집구조를 분석하고 국립공원의 식생경관관리를 위한 기초자료를 마련하고자 오대산 국립공원의 노인봉지역에 12개 조사구를 선정하고 군집구조분석을 실시하였다. TWINSPAN 분석결과 군집은 서어나무-신갈나무군집(군집 1), 거제수나무-박달나무-서어나무군집(군집2)으로 분리되었다. 군집의 천이단계는 종조성, 유사도지수, 종다양성분석 결과 극상림단계 초기와 극상림으로 밝혀졌다. 종수 ? 개체수분석에서는 100m$^{2}$당 평균 120~130개체, 17종이 조사되었다. 흉고단면적과 흉고직경급별 분석에서는 교목층에서 신갈나무를 대신하여 서어나무, 거제수나무, 박달나무 등이 극상수종이 될 것이며 아교목층에서는 당단풍이 우점종을 이룰 것으로 판단되었다.

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내장산 국립공원의 현존식생과 잠재자연식생 (Actual Vegetation and Potential Natural Vegetation of Naejangsan National Park, Southwestern Korea)

  • Kim, Jeong-Un;Yang-Jai Yim
    • The Korean Journal of Ecology
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    • 제11권3호
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    • pp.145-152
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    • 1988
  • The potential natural vegetation of Naejangsan national park area, southwestern Korea, was inferred from the actual vegetation. With the phytosociological classification, ordinatins and field surveys, the actual vegetation map of the area was made in scale 1:25, 000, including ten communities of Pinus densiflora, quercus mongolica, Quercus variabilis, Carpinus laxiflora, Daphnipyllum macropodum, Carpinus tschonoskii, Quercus aliena-Carpinus tschonoskii, Cornus controversa-Lindera erythrocarpa, Torreya mucifera-Zelkova serrate and Acer mono-Zelkova serrata community. The analyses of species richness, age structure and various informations on vegetation changes suggest the three pathways of late stage succession from P. densiflora forest to climatic climax. The first of them is through Q. variabilis forest to Q. monogolica forest in the upper parts of the mountain, the second through Q. variabilis and Q. serrata forest to C. laxiflora forest in the middle parts and the third through Q. aliena forest to C. tschonoskii forest in lower parts. Considering the actual vegetation and informations on the vegetation changes including human activities, the potential natural vegetation of the mountain mainly composed of Q. monogolica, C. laxiflora, C. tschonoskii, P. densiflora and Z. serrata forest as climatic climax and/or edaphic climax was inferred. The present situration of nature conservation in the area was estimated by the examination on the actual vegetation and potential natural vegetation map.

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Using Chemical and Biological Approaches to Predict Energy Values of Selected Forages Affected by Variety and Maturity Stage: Comparison of Three Approaches

  • Yu, P.;Christensen, D.A.;McKinnon, J.J.;Soita, H.W.
    • Asian-Australasian Journal of Animal Sciences
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    • 제17권2호
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    • pp.228-236
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    • 2004
  • Two varieties of alfalfa (Medicago sativa L cv. Pioneer and Beaver) and timothy (Phleum pratense L cv. Climax and Joliette), grown at different locations in Saskatchewan (Canada), were cut at three stages [1=one week before commercial cut (early bud for alfalfa; joint for timothy); 2=at commercial cut (late bud for alfalfa; pre-bloom head for timothy); 3=one week after commercial cut (early bloom for alfalfa; full head for timothy)]. The energy values of forages were determined using three approaches, including chemical (NRC 2001 formula) and biological approaches (standard in vitro and in situ assay). The objectives of this study were to determine the effects of forage variety and stage of maturity on energy values under the climate conditions of western Canada, and to investigate relationship between chemical (NRC 2001 formula) approach and biological approaches (in vitro and in situ assay) on prediction of energy values. The results showed that, in general, forage species (alfalfa vs. timothy) and cutting stage had profound impacts, but the varieties within each species (Pioneer vs. Beaver in alfalfa; Climax vs. Joliette in timothy) had minimal effects on energy values. As forage maturity increased, the energy contents behaved in a quadratic fashion, increasing at stage 2 and then significantly decreasing at stage 3. However, the prediction methods-chemical approach (NRC 2001 formula) and biological approaches (in vitro and in situ assay) had great influences on energy values. The highest predicted energy values were found by using the in situ approach, the lowest prediction value by using the NRC 2001 formula, and the intermediate values by the in vitro approach. The in situ results may be most accurate because it is closest to simulate animal condition. The energy values measured by biological approaches are not predictable by the chemical approach in this study, indicating that a refinement is needed in accurately predicting energy values.

Gap 모델을 이용한 광릉삼림군락의 천이에 대한 모의 실험 (Simulation of Forest Succession in Kwangnung Experimental Forest with Gap Model)

  • Han, You-Young;Park, Seung-Tai;Kim, Joon-Ho;Lee, Chang-Seok
    • The Korean Journal of Ecology
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    • 제19권6호
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    • pp.499-506
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    • 1996
  • Forest stand development in Kwangnung Experimental Forest, Korea, was simulated with a forest succession gap model of the JABOWA/FORET type, in order to predict climax species and characterime the trend of community structure along the succession. The model runs for a period or 1, 000 yr and is based on the averaged successional characteristics of 50 forest plote with an individual size or 1/12 ha gap consisted of the 15 major tree species. The total bimass and leafarea index have arrived at a steady state since about 200 yr and these values are smaller than that or field survey. Carpinus cordata, C. laxiflora, Quercus mongolica and Q. serrata were epected to be climax species that represent about 86% or total biomass in later stage and these results coincided with the previous succession studies from field survey in the area.

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생태형태학적(生態形態學的) 특성(特性) 분석(分析)에 의한 활엽수종(闊葉樹種)의 극성상지수(極盛相指數) 추정(推定) (The Estimation of Climax Index for Broadleaved Tree Species by Analysis of Ecomorphological Properties)

  • 김지홍
    • 한국산림과학회지
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    • 제82권2호
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    • pp.176-187
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    • 1993
  • 천연활엽수림(天然闊葉樹林) 군집(群集)에서 수종(樹種)의 천이계열(遷移系列)상의 위치를 해석(解析)하기 위하여 84개 활엽수(闊葉樹) 교목(喬木) 및 관목(灌木) 수종들의 생태형태학적(生態形態學的) 특성 분석을 바탕으로 극성상지수(極盛相指數)를 추정(推定)하였다. 생태형태학적(生態形態學的) 특성은 천이(遷移) 단계와 관계있다고 판단한 19가지를 선정하였으며, 각 수종별로 특성마다 극성상으로 갈수록 증가하는 2-4단계의 표준화된 점수를 부여하고 총점에 대한 합계점수의 백분율로써 극성상지수(極盛相指數)로 삼았다. 연구 대상 수종 중에서 서어나무의 지수(指數)가 83.3으로 최고치를 기록하였고 사시나무의 지수(指數)가 18.8로써 최저치로 추정되었으며, 전체 지수(指數)의 평균(平均)은 54.2로 산출되었다. 70이상의 지수(指數)값을 나타낸 수종은 9개, 40이상 70미만의 지수(指數)값을 나타낸 수종은 58개, 그리고 40미만의 지수(指數)값을 나타낸 수종은 17개로 집계(集計)되어, 천이(遷移) 중반단계의 삼림이 갖는 다양한 자원(資源) 혹은 생태적(生態的) 지위(地位)(niche)를 이용하는 수종(樹種)의 수가 압도적으로 많음을 알 수 있다. 주성분분석(主成分分析)을 통하여 각 수종이 광선흡수(光線吸收), 번식(繁殖), 그리고 목재(木材) 성질(性質) 등의 요인(要因)에 따르는 위치를 3차원(次元) 좌표(座標)상에 ordination하였고, cluster분석(分析)을 통하여 유사(類似)한 특징을 가진 4가지 수종군(樹種群)을 분류(分類)하였다. 과(科)별로 극성상지수(極盛相指數) 범위를 파악한 결과, 자작나무과(科)와 단풍나무과(科)에 속하는 수종들의 지수(指數) 범위가 넓었고, 버드나무과의 수종들은 선구수종의 전형적인 특성을 나타내었으나, 특별히 극성상(極盛相) 생활형(生活形)의 수종군(樹種群)을 갖는 과(科)는 없었다.

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