• Journal of Forest and Environmental Science
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• v.28 no.4
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• pp.199-204
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• 2012

This study was conducted to estimate climax index by eco-morphology for major 36 tree and sub-tree species in natural deciduous forests so as to interpret seral position of each species in the forest community. Fourteen eco-morphological characters which were considered to be associated with successional gradient in the forest were selected for the study. Four levels per character for each species were given on a standardized scale of increasing climax, and the index was computed by the proportion of the sum of total scores, expressed by percent values. With calculated mean value of 54.8 for all indices, Carpinus cordata had the highest index value of 90.5, and Populus davidiana recorded the lowest of 13.2. The most climax group, greater than 70 of the index, contained only 8 species, intermediate group, between 41 to 70 of the index, had 23 species, and the most pioneer group, less than 40 of the index comprised 5 species. The result has noticed that the large number of species would take advantage of most diverse resource and niche in the intermediate stage of the sere in the forest. By cluster analysis all 36 species were subjected to be classified into several species groups which had common similar eco-morphological characteristics. The indices were additionally plotted on the two dimensional graph to recognize the positions related to the light absorption factor and reproduction factor. The climax index of tree and sub-tree species developed by this study could be applied to understand the present status of successional stage on the basis of species composition by the method of summing up the indices. And comparison of forest successional stage among various forest communities could be done by summing up the climax indices of composed species in each community. However, this kind of applied methodology should be limited to the forest of similar species composition and site condition.

• Korean Journal of Environment and Ecology
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• v.32 no.1
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• pp.77-96
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• 2018

We investigated and analyzed three Korean island sites (Bijin-do, Ae-do, and Bogil-do) and one Japanese site (Tachibanayama) of sword-leaf litsea (Actinodaphne lancifolia) forests, known as the climax forest, to discuss the vegetation succession sere of warm-temperature evergreen broad-leaved forests. We then reviewed the literature in Korea, Japan, China, and Taiwan to consider the distribution characteristics of evergreen broad-leaved forests, vegetation succession sere, and climax tree species. Although Mt. Tachibana and Ae-do showed the most advanced vegetation structure, the soil and ordination (CCA) analysis indicated that it was not enough to consider that the sword-leaf litsea forest was at the climax stage in the warm-temperature region. The Actinodaphne lancifolia forest is sparsely distributed in Korea and Japan while the common types of vegetation in the warm temperate zone region in East Asia are Machilus spp., Castanopsis spp., and Cyclobalanopsis spp. The vegetation succession sere of the Korean warm-temperature region is thought to have a secondary succession such as Pinus thunbergii, P. densiflora, Q. serrata (early stage) through Machilus thunbergii, innamomum yabunikkei, Neolitsea sericea, Actinodaphne lancifolia (middle stage) to Castanopsis sieboldii, Q. acuta, Q. salicina (climax stage). However, Machilus thunbergii will be the climax species as an edaphic climax in places where there is a strong influence of the sea wind, or it is difficult to supply the seeds of Castanopsis spp. and Cyclobalanopsis spp.

• Korean Journal of Veterinary Research
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• v.33 no.2
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• pp.309-320
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• 1993

Guinea pigs were cutaneously inoculated with Trichophyton verrucosum var album, that is a common causative fungus of dermatophytosis in cattle. The developmental process of lesions, clinical and histopathological changes and reisolations of the fungi were studied to evaluate the pathogenicity of Trichophyton verrucosum var album in guinea pigs. Results obtained through the experiments were summarized as follows : 1. The incidence of infection of the clipping group was 13(86%) of 15 animals, and that of the plucking group was 14(93%) of 15 animals. In both of the clipping and plucking groups, visible cutaneoas lesions were developed between 4 and 7 days post inoculation(p.i.). The spreading and the climax stages persisted for 4 to 11 and 6 to 12 days, respectively. 2. In macroscopic observations, formation of various degree of erythemas and scales over the inoculated skin sites were observed in the spreading stage. In the climax stage, exudative changes and dark red crusts were formed as typical circular lesions. In the healing stage, the lesions revealed shedding of crust, alopecia and hair regrowth. 3. In histopathological observations, infiltration of inflammatory cells, hyperplasia, microabscesses and keratinous-hyaloid materials of epidermis were observed in the spreading stage. Hyphal invasion was primarily observed at the level of epidermis and pilosebaceous ducts. In the climax stage, the infected epidermis was thick with severe hyperplasia, hyperkeratosis and acanthosis. The microabscesses with fungal hyphae, folliculitis and hyperplasia of external root sheath were observed in the dermis. The fungal hyphae were observed only in the tissues of hair follicles, that were internal root sheath, cuticle, the keratinized portions of cortex and medulla 4. In reisolation of the inoculated fungus, all trials for ten animals showed positive cultures until 25 days p.i.. Afterward, the reisolation rates were gradually decreased, showing all negative after 40 days p.i..

• The Korean Journal of Ecology
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• v.20 no.6
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• pp.439-450
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• 1997

The potential natural vegtation of the Pukhansan National Park area, mid-western Korea, was inferred from the actual vegetation. With the phytosociological classification and field surveys, the actual vegetation map of the park area was made on a scale of 1:25, 000, including fourteen communities. By the analyses of the species diversity, the age structure, the human interferences and various informations on vegetation changes, two pathways of late stage succession from Pinus densiflora forests to the climatic climax were suggested. One is from Quercus serrata forests to Q. mongolica forests throughout the mountain and the other, from Q. variabilis or Q. acutissima forests to Carpinus laxiflora forests in lower parts. Considering the vegetation changes, the potential natural vegetation of the park area mainly composed of Q. mongolica, C. laxiflora, P. densiflora and Zelkova serrata forest as the climatic and/or edaphic climax was inferred.

• The Korean Journal of Ecology
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• v.16 no.4
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• pp.459-476
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• 1993

Vegetational structure and successional sere were investigated for the evergreen and nearby deciduous broad-leaved forests in inland areas (Turyunsan, Pulgapsan) and south- western island areas (Hongdo, Chindo). The evergreen broad-leaved forests of the four areas showed a sera1 stage with an age structure mostly under fifty years old. Dominant evergreen broad-leaved species in canopy layer were Castanopsis cuspidata var. sieboldii, Machilus thunbergii, Quercus acuta, Q. stenophylla, Q. glauca, and Ncolitsea serzceu. Succession of the forest vegetation in the surveyed areas was proceeding from deciduous broad-leaved trees to evergreen broad-leaved trees. The species of climax stage were assumed to be Machilus thunbergii, iVeolitseu sericea and Dendropanax morbifera in canopy layer, Aucuba japonica and Machilus japonica in subtree and shrub layer.

• Korean Journal of Environment and Ecology
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• v.9 no.2
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• pp.156-165
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• 1996

To investigate the forest structure and to suggest the management of vegetation landscape in Noinbong area, Pdaesan National Pa, twelve plots were set up and surveyed. According to the acalysis of classification by TWINSPAN, the community was divided by two groups of Carpinus laxiflora - Quercus mongolica community and the other is Betula costata - schmidtii - C. laxiflora community. It was found out that the successional stage of Noinbong forests was climax and introduced-climax by the analysis of species structure, similarity index and species diversity. The number of individuals was about 120~130 and species was 17 per 100m$^{2}$. Through the analysis of basal area and DBH class distribution, it was estimated that C. laxiflora, B. costata, and B. schmidtii will be clmax species instead of Q. mongolica in tree layer, and in the subtree layer, Acer pseudo-sieboldianum will be dominant species.

• The Korean Journal of Ecology
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• v.11 no.3
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• pp.145-152
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• 1988

The potential natural vegetation of Naejangsan national park area, southwestern Korea, was inferred from the actual vegetation. With the phytosociological classification, ordinatins and field surveys, the actual vegetation map of the area was made in scale 1:25, 000, including ten communities of Pinus densiflora, quercus mongolica, Quercus variabilis, Carpinus laxiflora, Daphnipyllum macropodum, Carpinus tschonoskii, Quercus aliena-Carpinus tschonoskii, Cornus controversa-Lindera erythrocarpa, Torreya mucifera-Zelkova serrate and Acer mono-Zelkova serrata community. The analyses of species richness, age structure and various informations on vegetation changes suggest the three pathways of late stage succession from P. densiflora forest to climatic climax. The first of them is through Q. variabilis forest to Q. monogolica forest in the upper parts of the mountain, the second through Q. variabilis and Q. serrata forest to C. laxiflora forest in the middle parts and the third through Q. aliena forest to C. tschonoskii forest in lower parts. Considering the actual vegetation and informations on the vegetation changes including human activities, the potential natural vegetation of the mountain mainly composed of Q. monogolica, C. laxiflora, C. tschonoskii, P. densiflora and Z. serrata forest as climatic climax and/or edaphic climax was inferred. The present situration of nature conservation in the area was estimated by the examination on the actual vegetation and potential natural vegetation map.

• Asian-Australasian Journal of Animal Sciences
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• v.17 no.2
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• pp.228-236
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• 2004

Two varieties of alfalfa (Medicago sativa L cv. Pioneer and Beaver) and timothy (Phleum pratense L cv. Climax and Joliette), grown at different locations in Saskatchewan (Canada), were cut at three stages [1=one week before commercial cut (early bud for alfalfa; joint for timothy); 2=at commercial cut (late bud for alfalfa; pre-bloom head for timothy); 3=one week after commercial cut (early bloom for alfalfa; full head for timothy)]. The energy values of forages were determined using three approaches, including chemical (NRC 2001 formula) and biological approaches (standard in vitro and in situ assay). The objectives of this study were to determine the effects of forage variety and stage of maturity on energy values under the climate conditions of western Canada, and to investigate relationship between chemical (NRC 2001 formula) approach and biological approaches (in vitro and in situ assay) on prediction of energy values. The results showed that, in general, forage species (alfalfa vs. timothy) and cutting stage had profound impacts, but the varieties within each species (Pioneer vs. Beaver in alfalfa; Climax vs. Joliette in timothy) had minimal effects on energy values. As forage maturity increased, the energy contents behaved in a quadratic fashion, increasing at stage 2 and then significantly decreasing at stage 3. However, the prediction methods-chemical approach (NRC 2001 formula) and biological approaches (in vitro and in situ assay) had great influences on energy values. The highest predicted energy values were found by using the in situ approach, the lowest prediction value by using the NRC 2001 formula, and the intermediate values by the in vitro approach. The in situ results may be most accurate because it is closest to simulate animal condition. The energy values measured by biological approaches are not predictable by the chemical approach in this study, indicating that a refinement is needed in accurately predicting energy values.

• The Korean Journal of Ecology
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• v.19 no.6
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• pp.499-506
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• 1996

Forest stand development in Kwangnung Experimental Forest, Korea, was simulated with a forest succession gap model of the JABOWA/FORET type, in order to predict climax species and characterime the trend of community structure along the succession. The model runs for a period or 1, 000 yr and is based on the averaged successional characteristics of 50 forest plote with an individual size or 1/12 ha gap consisted of the 15 major tree species. The total bimass and leafarea index have arrived at a steady state since about 200 yr and these values are smaller than that or field survey. Carpinus cordata, C. laxiflora, Quercus mongolica and Q. serrata were epected to be climax species that represent about 86% or total biomass in later stage and these results coincided with the previous succession studies from field survey in the area.

• Journal of Korean Society of Forest Science
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• v.82 no.2
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• pp.176-187
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• 1993

Based on the analysis of ecomorphological characters for 84 tree and shrub species, climax indices were estimated so as to interpret the position of the successional sere for the species in the natural deciduous forest. Nineteen ecomorphological characters, considered to be associated with successional gradient in the forest, were selected for the study. One of 2 to 4 steps per character for each species was given on a standardized scale of increasing climaxness, and the index was computed by percent of the sum of the scoring values for total score. Calculated mean value of 54.2 for all indices. Carpinus laxiflora had the highest index value of 83.3, and Populus davidiana recorded the lowest of 18.8. The most climax group, greater than 70 of the index, contained only 9 species, intermediate group, between 40 to 70 of the index, had 58 species, and the most pioneer group, less than 40 of the index comprised 17 species. The result has noticed that the large number of species would take advantage of most diverse resource and niche in the intermediate stage of the sere in the forest. The three components, i.e., light absorption, reproduction, and wood quality were used as axes for a 3-dimensional projection of the relative position for 44 species by principal component analysis. Along the similar ecomorphological characters, four recognized species group were classified by cluster analysis. The distribution pattern of plant families on the index gradient showed that the Betulaceae and Aceraceae had the widest seral amplitude, and the Salicaceae was a family typified as pioneer. There were no families specializing entirely with climax niche.