• 제목/요약/키워드: leaf variation

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Intraspecific Variation in Leaf Life Span for the Semi-evergreen Liana Akebia trifoliata is Caused by Both Seasonal and Aseasonal Factors in a Temperate Forest

  • Kohei, Koyama;Kikuzawa, Kihachiro
    • Journal of Ecology and Environment
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    • 제31권3호
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    • pp.207-211
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    • 2008
  • We investigated the leaf demography of a temperate woody liana, Akebia trifoliata, in a temperate forest in Japan, Akebia is semi-evergreen: some leaves are shed before winter, while others remain through the winter. Previous studies of semi-evergreen species found that variation in leaf life span was caused by variation in the timing of leaf emergence, Leaves that appeared just before winter over-wintered, while leaves appearing earlier were shed, However, it is unclear whether leaves of the same cohort (i.e., leaves that appear at the same time within a single site) show variation in life span under the effect of strong seasonality. To separate variation in life span among the leaves in each cohort from variation among cohorts, we propose a new method - the single leaf diagram, which shows the emergence and death of each leaf. Using single leaf diagrams, our study revealed that Akebia leaves within a cohort showed substantial variation in life span, with some over-wintering and some not. In addition, leaves on small ramets in the understory showed great variation in life span, while leaves on large ramets, which typically reach higher positions in the forest canopy, have shorter lives, As a result, small ramets were semi-evergreen, whereas large ramets were deciduous, The longer lives of leaves on small ramets can be interpreted as a shade-adaptive strategy in understory plants.

지역(地域), 죽림분별(竹林分別)에 따른 왕대의 엽형질(葉形質) 변이(變異) (The Variation of Some Leaf Characteristics in Phyllostachys bambusoides by Districts and Stands)

  • 홍한표;이강영
    • 한국산림과학회지
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    • 제68권1호
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    • pp.46-51
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    • 1985
  • 우리나라에서 유용죽종(有用竹種)으로 취급(取扱)되고 있는 왕대의 몇가지 엽형질(葉形質)에 대(對)하여 지역(地域)과 죽림분간(竹林分間)에 따른 변이정도(變異程度)를 구명(究明)하여 다음과 같은 결과(結果)를 얻었다. 엽형질(葉形質)에 있어서 이리지역(裡里地域)의 변이계수(變異係數)가 다른 지역(地域)에 비(比)하여 크게 나타났으며 죽림분간(竹林分間)의 변이계수(變異係數)에 있어서도 큰 차이(差異)를 나타냈다. 또한 이리지역(裡里地域)을 제외(除外)한 다른 지역간(地或間)에 있어서 엽형질(葉形質)의 평균치(平均値)는 차이(差異)가 없었으나 지역내(地域內)의 죽림분간(竹林分間)에 있어서는 대부분이 유의차(有意差)가 인정(認定)되었다. 엽폭(葉幅)/엽신장(葉身長)의 형상비(形狀比) 출현율(出現率)에 있어서 경주(慶州), 울산지역(蔚山地域)은 다른 지역(地域)에 비(比)하여 다소(多少) 분리(分離)되는 경향(傾向)이었다. 이상(以上)의 결과(結果)에서 왕대의 엽형질(葉形質)은 지역간(地域間)보다 지역내(地域內)의 죽림분간(竹林分間)에 따라 변이폭(變異幅)이 크게 나타나고 있음을 알 수 있었다.

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생산연도 및 지역별 본엽 2등 잎담배의 주요 화학성분 함량 변이 (The Variations of Some Chemical Constituents of Leaf Tobacco(Leaf, Grade 2) Produced in Various Growing Areas from 1999 to 2003 Crop Years)

  • 김상범;정기택;조수헌;복진영;정열영;이종률
    • 한국연초학회지
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    • 제26권1호
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    • pp.17-26
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    • 2004
  • This study was conducted to get the informations for reducing the variation of chemical contents of leaf tobacco. The contents and variations of some chemical constituents of leaf(Leaf, Grade 2) produced in various growing areas from 1999 to 2003 and the effects of meteorological factors on the chemical constituents of leaf were analysed. The contents of analysed constituents of leaf showed high significant differences among crop years in flue-cured and burley, particularly the variation among crop years were higher in chlorine and nicotine contents while lower in total nitrogen content. There were significant differences among growing areas in nicotine and total sugar contents of flue-cured leaf and chlorine content of burley leaf. The total sugar content were negatively correlated to the nicotine and total nitrogen contents in flue-cured leaf. The average air temperature in June and July were positively correlated to the nicotine content of leaf while negatively to total sugar, and the precipitation in May were negatively correlated to the nicotine while positively to total sugar.

Seasonal Variation in Contents of Sugars in Different Parts of Broccoli

  • Bhandari, Shiva Ram;Kwak, Jung-Ho
    • 원예과학기술지
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    • 제33권2호
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    • pp.276-282
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    • 2015
  • Seasonal variation in the contents of sugars (fructose, glucose, and sucrose) in the floret, leaf, and stem of broccoli were studied in ten commercial broccoli cultivars. Plants were grown in the spring and fall seasons in 2011. In both seasons, glucose was the major constituent, comprising about 50% of the total sugar content in the floret and leaf tissue of most cultivars, whereas the broccoli stem showed an unusual pattern of accumulation. Sucrose exhibited greater cultivar dependency as well as seasonal variation compared to fructose and glucose in floret and leaf tissues. The floret tissue had a higher total content of sugar in the spring compared to the fall due to an increase in glucose and fructose. However, most of the leaf and stem tissues of broccoli had a higher total sugar content in the fall compared to the spring. Furthermore, stem and leaf tissues possessed a relatively higher total sugar content compared to floret tissue in both seasons. 'Grandeur' broccoli contained a significantly greater amount of total sugar in both floret and leaf tissues in both seasons, whereas 'YuDoRi No.1' broccoli exhibited the highest total content of sugar in stem tissue. At overall, the results showed significant influences of genotype, plant part and growing season on sugar content in broccoli.

Linking Leaf Functional Traits with Plant Resource Utilization Strategy in an Evergreen Scrub Species Rhododendron caucasicum Pall. along Longitudinal Gradient in Georgia (The South Caucasus)

  • Ekhvaia, Jana;Bakhia, Arsena;Asanidze, Zezva;Beltadze, Tornike;Abdaladze, Otar
    • Journal of Forest and Environmental Science
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    • 제38권2호
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    • pp.110-121
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    • 2022
  • Leaf functional traits widely have been used to understand the environmental controls of resource utilization strategy of plants along the environmental gradients. By using key leaf functional traits, we quantified the relationships between leaf traits and local climate throughout the distributional range of Rhododendron caucasicum Pall. in eastern and western Georgian mountains (the South Caucasus). Our results revealed, that all traits showed high levels of intraspecific variability across study locations and confirmed a strong phenotypic differentiation of leaf functional variation along the east-west longitudinal gradient in response to the local climate; out of the explored climatic variables, the moisture factors related to precipitation and number of precipitation and dry days for winter and growth seasons were more strongly related to leaf trait variation than the elevation and air temperature. Among studied leaf traits, the leaf specific area (SLA) showed the highest level of variability indicating the different resource utilization strategies of eastern and western-central Rh. caucasicum individuals. High SLA leaves for western-central Caucasian individuals work in relatively resource-rich environments (more humid in terms of precipitation amount and the number of precipitation days in winter) and could be explained by preferential allocation to photosynthesis and growth, while eastern Caucasian samples work in resource-poor environments (less humid in terms of precipitation amount and the number of precipitation days in winter) and the retention of captured resources is a higher priority appearing in a low SLA leaves. However, more evidence from a broader study of the species throughout its distribution range by including additional environmental factors and molecular markers are needed for firmer conclusions of intraspecific variability of Rh. caucasicum.

한국산 제비꽃속의 종분화론적 연구 - 태백제비꽃군의 비교형태학적 연구 - (Biosystematic Study on the Genus Viola in Korea - Comparative Morphology of the Viola albida Complex -)

  • 김경식
    • Journal of Plant Biology
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    • 제34권3호
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    • pp.229-238
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    • 1991
  • The main objective of this study was to clarify the taxonomic status and the speciation mechanism of Viola albida complex in Korea. The complex includes three very closely related taxa, taxonomic ranks of which are much in dispute. These taxa in the complex are sympatric in distribution, and they are very similar in most morphological characteristics. Comparative morphological analyses revealed that they can be distinguished on the basis of leaf shape, although they show complicated patterns of variation in leaf shape. The variation of leaf shape in each taxon appeared to be maintained through developmental process, and extensive cleistogamy and vegetative reproduction in these taxa are suggested as the mechanisms maintaining the variation. Genetic variation in this complex was monomorphic in all 14 loci examined. Palynological study also showed that these taxa have the same pollen morphology. These results strongly suggested that they should be recognized as varieties of V. albida; var, albida, var. chaerophylloides, and var. takahashii. The results also suggested that some individuals that can be assigned to var. takahashii might be hybrids between var. albida and var. chaerophylloides.

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Zoysiagrass 수집계통들과 종간교배계통들의 형태적 특성들의 변이 (Variation of the Morphological Characteristics in the Accessions of Zoysia Species and Their Hybrid Lines)

  • 김형기;김기선;주영규;홍규현;김경남;이재필;모숙연;김두환
    • 아시안잔디학회지
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    • 제10권1호
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    • pp.1-11
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    • 1996
  • Fifty accessions in Zoysia species and forty-one hybrid lines were evaluated in order to detect analyze the genectic variation of Zoysiagrasses and to obtain the basic information for breeding. 1.Plant height ranged from 13 to 34.5cm, leaf length from 7 to 26cm, leaf width from 2 to 7mm, and first leaf height from 1.1 to 8.5cm 2. The correlations among plant height, leaf length and first-leaf height were high, but leaf Width was correlated only With leaf length. 3. Stolon length was varied from 10 to 108cm, number of stolons from 12 to 53, internode length from 2 to 6.1cm and coverage percentage from 10 to 90%. 4.There were correlation among stolen length, number of stolons, internode length and coverage percentage except between stolen and internode length. 5.M1J and MC2 showed the highest level in stolen length, number of stolons and coverage percentage. Therefore they will be the best breeding materials for the purpose of fast covering. 6.According to the analysis on October 26, 1995, yellow color-change rate of leaves ranged from 5 to 85%, red color-change from 0 to 75% and sum of yellow and red change from 20 to 95%. Speciallty M2M1 and FL-41 kept green color untill early December, indicating that they had the late dormancy. 7. Visual assessment of stolon length and number of stolons, showed similar tendency With actually measured data. 8. Shoot density was very high in M2M1 and S2, and showed negative correlation with leaf width. 9. It is known in Korea that the darker green color of lawngrass is, the better it looks. There were 11 lines with dark green color leaves such as J 85-10, Meyer in Z. japonira, one asacession in Z. matrella and M2M1, M2S8, JM2, 55C2 in hybrid lines.

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희귀수종 먹넌출 엽의 형태적 특성과 유전변이 (Characteristics of Leaf Morphology and Genetic Variation of the Rare Woody Plant, Berchemia racemosa var. magna)

  • 송정호;임효인;장경환;한진규
    • 한국자원식물학회지
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    • 제26권5호
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    • pp.613-618
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    • 2013
  • 우리나라에서 안면도에만 분포하는 먹넌출 집단의 생명자원보존을 위하여 엽의 형태적 특성과 I-SSR 표지자를 이용한 유전변이를 조사하였다. 10가지 엽특성에 대한 ANOVA 분석결과 모든 특성에서 개체 간에 통계적인 유의성이 인정되었다. 조사된 39개체의 평균특성은 엽신장 11.8 cm, 최대엽폭 7.1 cm, 엽지수 1.67, 상1/3폭 5.4 cm, 하1/3폭 6.2 cm, 엽병길이 3.6 cm, 엽두께 0.19 mm, 엽맥수(좌) 11.5개, 엽맥수(우) 11.4개, 엽면적 61.7 $cm^2$로 나타났다. 변이계수 값은 엽두께, 엽병길이, 엽면적이 각각 18.8%, 21.7%, 22.0%로 높게 나타났으며, 나머지 특성들에서는 15% 이내의 비교적 낮은 변이를 나타냈다. 선발된 8개 I-SSR Primer에서 총 50개의 증폭산물을 얻었으며, 유효대립 유전자의 수 1.719개, 다형적 유전자좌의 비율 26.0%, 이형접합도의 기대치 0.410 및 Shannon의 다양성지수 0.598로 각각 나타났다. 안면도 먹넌출 집단은 제한된 지역에 분포하며 개체수가 적음에도 불구하고 높은 유전다양성을 유지하는 것으로 나타났다.

후박나무 10개 천연집단의 엽형질 변이 (The Leaf Morphological Variation of Ten Regions of Natural Populations of Machilus thunbergii in Korea)

  • 양병훈;송정호;이재천;박용구
    • 농업생명과학연구
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    • 제45권3호
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    • pp.25-33
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    • 2011
  • 후박나무의 유전자원보존을 위하여 10개 천연집단을 대상으로 9가지 엽의 형태적 특성에 대한 집단간 및 집단내 개체간 변이를 조사하고 다변량분석을 실시하였다. 엽의 평균 생장은 엽신장 9.8cm, 최대엽폭 4.0cm, 엽병길이 1.8cm, 엽맥수 8.4개, 엽저각 $67.9^{\circ}$, 엽두각 $78.0^{\circ}$로 나타났다. 각 형질특성에 대한 변이계수 값은 대체적으로 20% 내외의 비교적 유사한 특성을 나타냈다. Nested 분산분석 결과 모든 특성에서 집단간 및 개체간에 고도의 유의적인 차이가 인정되었으며, 전체 분산 가운데 집단간보다 집단내 개체간 차지하는 비율이 모든 특성들에서 높게 나타났다. 집단간 유연관계는 Euclidean distance 1.2 수준에서 크게 4개의 그룹으로 나뉘었으나 지리적 분포에 따른 특별한 경향은 나타나지 않았다. 유집군의 유형에 대한 주성분분석 결과 제3주성분까지가 누적변이 값이 92.8%로 나타났다. 제1주성분의 기여율은 40.3%로 대체적으로 최대엽폭, 제2주성분의 기여율은 28.7%로 엽신장, 제3주성분의 기여율은 23.8%로 엽병길이 특성의 기여도가 높게 나타나 후박나무 집단간 유연관계에 중요한 정보를 주는 요인으로 나타났다.

쑥에 함유된 monoterpenes 의 함량과 조성의 계절적 변이 (Seasonal Variation in Concentration and Composition of Monoterpenes from Artemisia princeps var. orientalis)

  • Kim, Jong-Hee
    • The Korean Journal of Ecology
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    • 제19권4호
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    • pp.321-328
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    • 1996
  • The profile and concentration of monoterpene metabolites in the leaf and stem of Artemisia princeps var. orientalis were quantified, and seasonal variation in monoterpenes of Artemisia plant was investigated. Samples were taken from five sites at the campus of Kyungnam University during maturing season. Monoterpenes in leaf and stem were analyzed using gas chromatography-mass spectrometry (GC-MS). The major constituents of A. princeps var. orientalis in both the leaf and stem were 21 monoterpenes.$ {\alpha}-pinene,\;{\beta}-pinene,\;{\beta}-myrcene$, dl-limonene, naphthalene and unknown monoterpenes with 5.49 and 16.27 of retenstion time were present in high concentrations of compounds identified on the leaf and stem of A. princeps var. orientalis. The cmounts of total monoterpenes of leaf were from two to five times higher than stem and rapidly decreased with the time, while that of stem was constnat except early spring. Most of the high percentage of monoterpenes in the leaf were those with later retention time. These results indicated that monoterpenes yields are considered to be more variable than monoterpene composition in responding to the time in both the leaf and stem.

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