• Title/Summary/Keyword: Shade tolerance

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Studies on the Shade Tolerance, Light Requirement and Water Relations of Economic Tree Species(II) -Effects of Artificial Shade Treatment on the Water Potential and Stomatal Diffusive Resistance of Four Deciduous Hardwood Species- (주요경제수종(主要經濟樹種)의 내음성(耐陰性) 및 광선요구도(光線要求度)와 수분특성(水分特性)에 관한 연구(硏究)(II) -인공피음(人工被陰)이 활엽수(闊葉樹) 4종(種)의 수분(水分)포텐셜 및 기공증산저항(氣孔蒸散低抗)에 미치는 영향(影響)-)

  • Kwon, Ki Won;Choi, Jeong Ho;Chung, Jin Cheol
    • Journal of Korean Society of Forest Science
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    • v.89 no.2
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    • pp.198-207
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    • 2000
  • The diurnal and seasonal changes of water potential and stomatal diffusive resistance were investigated with the effects of shade treatment to elucidate the water relations of the one year old seedlings of Betula platyphylla var. japonica, Zelkova serrata, Acer mono, and Prunus sargentii subjected to five levels of artificial shade treatments from full sun to nearly full shading of 2-6% relative light transmittance. Stomatal diffusive resistance measured in the four deciduous hardwood species studied changed variously by growing season and by species with the five shade treatments in the ranges of 1.7~25.1s/cm in 9 a.m., 2.3~33.3s/cm in 1 p.m., and 1.1~36.8s/cm in 5 p.m.. The measurements of stomatal diffusive resistance increased with increasing the shading, and were higher in September than in June or July in most of the species studied. The stomatal diffusive resistance measured in Zelkova serrata, Acer mono, and Prunus sargentii seemed to be approximately 3~4 times higher in nearly full shading treatment than in full sun. Water potential also changed variously by growing season and by species with five different shade treatments in the ranges of -0.17~-1.20MPa in 9 a. m., -0.30~-2.03MPa in 1 p. m., and -0.18~-1.34MPa in 5 p.m., respectively. On the reverse of stomatal diffusive resistance, the measurements of water potential were lower in September than in June or July in most of the species studied, and the seasonal differences were especially greater in Zelkova serrata comparing with the other species. The water potential seemed to be higher of approximately 2~3 times in nearly full shading treatment than in full sun in all of the species studied, but the differences among the shading treatments were less in water potential than in stomatal diffusive resistance. The differences of water potential following the gradient of five shading treatment seemed to be less in Betula platyphylla var. japonica than in the other species. In Acer mono and Prunus sargentii being some shade tolerant species, the water potential increased rapidly through about noontime with raising the shading level from full sun to the intermediate shading level of 22-28% relative light transmittance, but slowly with closing to full shading. In Betula platyphylla var. japonica and Zelkova serrata being shade intolerant species, the water potential increased gradually throughout the shading levels and the increment ranges were greater in Zelkova serrata than in Betula platyphylla var. japonica.

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Community Ecological Revaluation of Acer pseudosieboldianum and Carpinus cordata in the Natural Deciduous Forest

  • Kim, Ji Hong;Kang, Sung Kee;Lim, Seon Mi
    • Journal of Forest and Environmental Science
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    • v.32 no.1
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    • pp.74-81
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    • 2016
  • Classified into sub-tree of the growth-form, Acer pseudosieboldianum and Carpinus cordata hardly reach the uppermost canopy with limited height growth potentiality in the natural deciduous forest. However, the two species usually dominate lower stories of the forest, playing an important role in overall ecological characteristics in the forest. Based on the vegetation data of 106 $20m{\times}20m$ sample plots in Mt. Jumbong area, this study was carried out to evaluate ecological constitution by several quantitative analysis so as to understand the mechanism of the natural deciduous forest. The results indicated that individuals of A. pseudosieboldianum and C. cordata were absent or few in overstory, but emerged the most dominant species in midstory and understory, providing adequate proof of the ecological importance. The comparison of indices of succession between presented and predicted values in midstory did not make much difference, suggesting that the species composition would not change much and come close to steady state in midstory and understory. The pair combination of species association noted that A. pseudosieboldianum had significant positive association with C. cordata, Quercus mongolica, and Tilia amurensis had significant positive association with A. pseudosieboldianum, A. pictum subsp. mono and Fraxinus mandshurica but negative association with F. rhynchophylla. Being compared with other major canopy tree species in the study forest, the target species of A. pseudosieboldianum and C. cordata had strong regeneration strategies, partially characterized by large number of saplings and pole sized trees and high ratio of live crown, which indicated high shade tolerance to survive in the limited amount of light under the canopy. Even though A. pseudosieboldianum and C. cordata do not reach and occupy the canopy layer mainly due to the inherent growth form, they would have highest competitive potentiality to prosper and dominate in the midstory of the natural deciduous forest.

Fundamental Studies on the Landscape Use of Evergreen Ground Cover Plants in KANG WON Area -Propagation, Shade Tolerance, Cold Resistance, and Growth Rate (강원도 지역에서의 상록지피식물의 조경적 이용에 관한 기초적 연구(I) -증식, 내음성, 내한성, 생육량의 차이에 관하여)

  • 홍종운;이기철;허범람;원경렬;임병춘
    • Asian Journal of Turfgrass Science
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    • v.7 no.1
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    • pp.19-30
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    • 1993
  • Experiments were conducted to investigate propagation, shade tolerance, cold resistance and growth rate of evergreen ground cover plants ; Vinca minor K , Ajuga reptans L., Ophiopogon japonicus ker. f nanus hort , and Hedera helix.. The results were as follows : 1.It is convenient to use the growing case for rice seed to grow and transplant Vinca minor seed-ling. The most proper density of transplanting Vinea was 180 plants per 1m$^2$. At the end of growing season, the coverage came up to 90% The rooting ability of Hedera helix was best to cut from April to May (temperature 15~23˚C). The seedling length of Parthenocissus quinguefohlia were irregular and ranged from 5 cm of 200cm. 2. Ajuga reptans L., Ophiopogon j. and Vinca minor L. grew better under 50% light intensity than full sunlight. Particularly, under full sunlight and aestival high temperature, there were a few withering plants in Ajuga reptans L. , and V Vilica minor L. 3. Of Vinca minor K., Ajuga reptans L. , Ophiopogon japonicus ker. f. nanus hort. and Hedera helix. acclimatized in open field, where temperature was from -7~8˚C. There were few plants had damage in low temperature incubator to -16˚C after field acclimatization. 4.Tendrils of Parthenocissus quinguefolia strongly clung to the wire netting stone fence but intruded into the gap and cranny of the precast concrete fence and so clung to, so that it needed to install the bending net on the precast concrete fence.

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Leaf Growth of Seven Fruit Trees in Response to Different Lights for Garden Tree

  • Nam, Yu Kyeong;Lee, Jin Hee;Kwon, O Man
    • Journal of Environmental Science International
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    • v.27 no.10
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    • pp.875-881
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    • 2018
  • To select fruit trees suitable for shaded urban garden with Elaeagnus umbellata var. coreana, Malus domestica 'Alps Otome', Malus domestica 'Fujii', Prunus mume, Prunus persica for. persica, Rubus fructicosus, and Vaccinium corymbosum 'Reka', leaf growth in response to different light was investigated two years after 35% shade treatment. Leaf area of E. umbellata var. coreana, M. domestica 'Alps Otome', P. mume, P. persica for. persica, and V. corymbosum 'Reka' increased in shading. Fresh weight of leaves make inconsistent response to shading in every species but dry weight of E. umbellata var. coreana and P. persica for. persica showed the highest 150% and 148%, increment, respectively. Althought leaf water content of E. umbellata var. coreana decreased in shading, there is no difference in P. persica for. persica. Chlorophyll value of E. umbellata var. coreana and P. persica for. persica that showed higher than any other species is correlated with dark leaf green. Compared to specific leaf weight of E. umbellata var. coreana, P. persica for. persica, and V. corymbosum 'Reka' showed lower than any other species in shading, that of M. domestica 'Fujii', and R. fructicosus increased in reverse. These results indicate that E. umbellata var. coreana and P. persica for. persica that showed high value in several invesetigaton items are suitable for shady urban condition considering leaf growth in response to shading.

Present and Future Prospect in Zoysia Grass Breeding (한국잔디류(Zoysia Grass)의 육종현황)

  • 주영규;김두환;이재필;모숙연
    • Asian Journal of Turfgrass Science
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    • v.11 no.1
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    • pp.73-85
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    • 1997
  • Although zoysia grass (Zoysia spp.) has a lot of excellent chracteristics as warm-season grass, it have been limited in use due to slow establishment, low seed production, poor shade tolerance and other factors. Breeding trials have been continued from 1900's, much attentions have been paid especially in U.S.A., Korea and Japan recently. In U.S.A., more than 24 varieties had been evaluated at National Turfgrass Evaluation Program(NTEP) from 1991 to 1995 and some were regsistered as commercial. After the 6th International Turfgrass Research Conference at Japan in 1989, Japan Turfgrass In-corporation (JTI) sponsored by private companies and government carried out breeding programs for pest, salt, and shade tolerant and herbicide resistant varieties. JTI also has been trying to im-prove vigor and breed evergreen zoysia Korean breeders collected germplasms since 1960's. After USDA breeders came to Korean penesula in 1982, Korean breeders joined with USDA zoysia breeding project for several years. Many interspecific hybrids and natural selected varieties were breeded that period both in U.S.A and Korea. Breeding objectives were to extend green color period, improve leaf quality and density, and better leaf color at dormant stage. Since 1990's, zoysia grass breeding trials are getting more diverse in many points such as random amplified polymorphic DNA (RAPD) assay for ecotype identification. The objectives of this study are to evaluate germplasms in Korea, and also review the present status and future prospect in zoysia grass breeding in the world.

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Photosynthetic Responses and Growth Performances in the Three Deciduous Hardwood Species Under Different Shade Treatments (피음처리를 실시한 3개 활엽수종의 광합성과 생장 특성)

  • Cho, Min Seok;Kwon, Ki Won;Choi, Jeong Ho
    • Journal of Korean Society of Forest Science
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    • v.96 no.4
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    • pp.462-469
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    • 2007
  • The present study was conducted to investigate photosynthetic responses and growth performances of Liriodendron tulipifera, Betula costata, and Carpinus cordata seedlings growing under four different light regimes (full sun and 54~65%, 26~37%, 8~13% of full sun). L. tulipifera and B. costata showed the highest photosynthesis rate of 13.59, $16.29{\mu}mol\;CO_2{\cdot}m^{-2}{\cdot}s^{-1}$, respectively, at full sun. And photosynthesis ability and growth performances were decreased as increasing shade level in the above two species. C. cordata showed the highest photosynthesis rate of $9.47{\mu}mol\;CO_2{\cdot}m^{-2}{\cdot}s^{-1}$ at 26~37% of full sun. Also, growth performances of height, root collar diameter, and biomass showed the results similar to photosynthetic responses presented in C. cordata.

An Analysis of Permanantly Shaded Areas and the Defect Rate of Landscape Trees in Apartment Complexes Using Daylight Simulations

  • Park, Sang Wook
    • Journal of People, Plants, and Environment
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    • v.23 no.3
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    • pp.333-345
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    • 2020
  • Background and objective: The purpose of this study was to provide basic data on trees that can be used for planting design and construction for permanently shaded areas by grasping the growth status of trees according to the daylight conditions of the outdoor spaces of apartment complexes. Methods: On the recently completed apartment complexes, daylight conditions were analyzed by using daylight simulations utilizing Solar Access Analysis of Ecotect Analysis. With a criteria for assessment of tree condition, the defect rate of trees planted in permanently shaded areas and green spaces with good daylight conditions was analyzed to suggest trees applicable to permanently shaded areas. The first tree survey was conducted from November 18, 2019 to February 15, 2020, focusing on trees planted in permanently shaded areas, and the second tree survey of all the trees planted on the study sites including permanently shaded areas was conducted from March 16 to March 30, 2020. Results: Evergreen trees which are classified as shade intolerant trees including Pinus densiflora, Thuja occidentalis, and Abies holophylla showed a higher defect rate of trees among the trees planted in permanently shaded areas. Taxus cuspidata, Zelkova serrata, Cornus kousa, Chionanthus retusus and Acer palmatum which are classified as shade tolerant trees and shade moderate tolerance trees seemed to be able to be used in the plant design of permanently shaded areas in apartment complexes because the trees showed good growth and a low tree defect rate. In addition, although it was excluded from the analysis due to a small number of samples, Sorbus commixta and Prunus cerasifera var. atropurpurea also can be used for planting in permanently shaded areas. Conclusion: The daylight simulation technique used to analyze permanent shaded areas in this study can be used as an analysis tool considering the daylight environment at the stages of design and construction, and additional research will be required to analyze tree growth according to daylight conditions through data accumulation and monitoring by managing records throughout the entire life cycle of trees in the process of planting and maintenance.

A Study on the Actual Conditionis and Improvement Plan of Interior Landscape in the Subway Stations (지하철역내의 실내조경현황과 개선방안에 관한 연구)

  • 방광자;주진희;박혜경;박성은
    • Journal of the Korean Institute of Landscape Architecture
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    • v.26 no.3
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    • pp.258-266
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    • 1998
  • The purpose of this study is to investigate the actual conditions, to find problems, and to present an improvement plan of interior landscape in the subway stationis. Moreoiver, it will presume the possibilties to develop the subway and ways to utilie underground space effectively. The results are summarized as follows A light intensity was recorded 0-100 Lux as the lowest light and 500 Lux below as the highest in 61 subway stations. An average temperature of 12 $^{\circ}C$and an air humidity of 38% were recorded over a 4 month period from January to April, but includes a drastic variaton between -8.5$^{\circ}C$ and 21.5$^{\circ}C$. Soil acidity of pH 7 below and soil moisture of 1-2 wet degree were apeared in subway stations mostly. Plant forms consisted of artificial flower & flower. Plant species were recorded at a total of 54 species and appeared in the order of Araucaria heteropylla, Ophiophogon jaburan, Aspidistra elatior cv. 'Asahi' and Hedera helix. Plant height was, for the most part, below 0.5m. Plant species that was fined of conditions were Palm, Camellia japonica, Araucaria heterophylla as a high plant, dracaena fragrans, dracaena deremensis cv. Wakneckii as a middle plant, and Ophiopogon jaburan, Hedra helix, ytomium falcatum, Aspidistra elatior cv. Asahi as a low plant. It used to water materials such as small pool, small cascade, water cycles and natural materials such as natural rock, small rock, sand, bark and animal materials such as squirrels, birds, goldfish as an object for plants in the subway stations. From these actual conditions, First of all, It must make up physical environments such as light, temperature humidity, soil for plant growth, and is important to chooce suitable indoor plants and draw up systematic management in the subway environments. Also, it change plants frequently and uses variable objects for subway stationi individuatism, Moreover, indoor plants with strong environmental adaptation abities such as shade tolerance, drought tolerance and cold tolerance need to develope variable species possibly. If these improvements occur, utilization and amenity of subway stationis will increase, according to the use of interior landscape.

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Photochemical Index Analysis on Different Shading Level of Garden Plants (정원 식물의 차광 조건별 광화학적 생리지표 해석)

  • Kang, Hong Gyu;Kim, Tae Seong;Park, So Hyun;Kim, Tae Wan;Yoo, Sung Yung
    • Korean Journal of Environmental Biology
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    • v.34 no.4
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    • pp.264-271
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    • 2016
  • The objective of this study was to determine the growth and light utilization efficiency of garden plants in shade area through chlorophyll fluorescence reaction analysis. Ten garden plants was grown for 75 days under 50% and 80% shading conditions. Under shading, $ET2_O/RC$, the fluorescence parameter related to electron-transport in photosystem II, was effectively enhanced. However, the electron transport flux until PSI acceptors per reaction center ($RE1_O/RC$) was reduced. These changes in photochemical parameters evoked a decrease in performance index (PI) and driving force (DF) of electron transport flux. In addition, some photochemical parameters such as $F_V$, $FV/F_O$, $RE1_O/RC$, $ET2_O/RC$, $PI_{TOTAL\;ABS}$, and $DF_{TOTAL\;ABS}$ were found to be important for shade tolerance. Three species (Pachysandra terminalis Siebold & Zucc, Physostegia virginiana L., and Carex maculata Bott) were found to be shade tolerant. Based on these results, shading factor index (SFI) deduced from photochemical parameters is useful for evaluating of shading stress of garden plants.

Disturbance regime and tree regeneration in kwangnung natural forest (광릉 자연림에서의 교란체제와 수목의 재생)

  • Cho, Do-Soon
    • The Korean Journal of Ecology
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    • v.15 no.4
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    • pp.395-410
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    • 1992
  • Disturbance regime and tree regeneration were studied in kwangnung natural forest, an old-growth deciduous hardwood forest located in central korea. This forest is dominated by carpinus laxiflora, c.erosa, and quercus species. The area occupied by canopy gaps was 4.6% of the total forested area, and the mean size of canopy gaps was 92 $m^2$ with the maximum being 524 $m^2$. More than half of the gaps were less than four years old, and 3/4 of the gaps were created by death of only or two canopy trees, indicating the dominance of small-sized gaps in kwangnung forest. about half of the gap-makers were c. laxiflora, and another one third were quercus species. In contrast, the most frequent relacers were c. laxiflora while quercus species filled only 5% of the gaps, suggesting a future shift in tree species composition under the current disturbance regime. tree regeration was more conspicuous even in small gaps than non-gaps regardless of shade-tolerance of tree species, indicating the importance of gaps in tree regeneration.

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