• Title/Summary/Keyword: LEAF VARIATION

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Intraspecific Variation in Leaf Life Span for the Semi-evergreen Liana Akebia trifoliata is Caused by Both Seasonal and Aseasonal Factors in a Temperate Forest

  • Kohei, Koyama;Kikuzawa, Kihachiro
    • Journal of Ecology and Environment
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    • v.31 no.3
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    • pp.207-211
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    • 2008
  • We investigated the leaf demography of a temperate woody liana, Akebia trifoliata, in a temperate forest in Japan, Akebia is semi-evergreen: some leaves are shed before winter, while others remain through the winter. Previous studies of semi-evergreen species found that variation in leaf life span was caused by variation in the timing of leaf emergence, Leaves that appeared just before winter over-wintered, while leaves appearing earlier were shed, However, it is unclear whether leaves of the same cohort (i.e., leaves that appear at the same time within a single site) show variation in life span under the effect of strong seasonality. To separate variation in life span among the leaves in each cohort from variation among cohorts, we propose a new method - the single leaf diagram, which shows the emergence and death of each leaf. Using single leaf diagrams, our study revealed that Akebia leaves within a cohort showed substantial variation in life span, with some over-wintering and some not. In addition, leaves on small ramets in the understory showed great variation in life span, while leaves on large ramets, which typically reach higher positions in the forest canopy, have shorter lives, As a result, small ramets were semi-evergreen, whereas large ramets were deciduous, The longer lives of leaves on small ramets can be interpreted as a shade-adaptive strategy in understory plants.

The Variation of Some Leaf Characteristics in Phyllostachys bambusoides by Districts and Stands (지역(地域), 죽림분별(竹林分別)에 따른 왕대의 엽형질(葉形質) 변이(變異))

  • Hong, Han Pyo;Lee, Kang Young
    • Journal of Korean Society of Forest Science
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    • v.68 no.1
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    • pp.46-51
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    • 1985
  • This study was carried out to know the variation of some leaf characteristics in Phyllostachys bambusoides by district and stands in Korea. The results obtained were summarized as follows ; The coefficients of variation of leaf characteristics in Iri were higher than the other districts. The variation coefficients of leaf characteristics were highly shown among stands in districts. The leaf characteristics were not significant differences in districts except in Iri. The leaf characteristics among stands were significant differences. The frequency distributions of form quotient of leaf blade width/leaf blade length were shown to be different from Gyeongju and Ulsan districts. In conclusion, the variations of leaf characteristics of Phyllostachys bambusoides were highly showed among stands in districs.

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The Variations of Some Chemical Constituents of Leaf Tobacco(Leaf, Grade 2) Produced in Various Growing Areas from 1999 to 2003 Crop Years (생산연도 및 지역별 본엽 2등 잎담배의 주요 화학성분 함량 변이)

  • 김상범;정기택;조수헌;복진영;정열영;이종률
    • Journal of the Korean Society of Tobacco Science
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    • v.26 no.1
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    • pp.17-26
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    • 2004
  • This study was conducted to get the informations for reducing the variation of chemical contents of leaf tobacco. The contents and variations of some chemical constituents of leaf(Leaf, Grade 2) produced in various growing areas from 1999 to 2003 and the effects of meteorological factors on the chemical constituents of leaf were analysed. The contents of analysed constituents of leaf showed high significant differences among crop years in flue-cured and burley, particularly the variation among crop years were higher in chlorine and nicotine contents while lower in total nitrogen content. There were significant differences among growing areas in nicotine and total sugar contents of flue-cured leaf and chlorine content of burley leaf. The total sugar content were negatively correlated to the nicotine and total nitrogen contents in flue-cured leaf. The average air temperature in June and July were positively correlated to the nicotine content of leaf while negatively to total sugar, and the precipitation in May were negatively correlated to the nicotine while positively to total sugar.

Seasonal Variation in Contents of Sugars in Different Parts of Broccoli

  • Bhandari, Shiva Ram;Kwak, Jung-Ho
    • Horticultural Science & Technology
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    • v.33 no.2
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    • pp.276-282
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    • 2015
  • Seasonal variation in the contents of sugars (fructose, glucose, and sucrose) in the floret, leaf, and stem of broccoli were studied in ten commercial broccoli cultivars. Plants were grown in the spring and fall seasons in 2011. In both seasons, glucose was the major constituent, comprising about 50% of the total sugar content in the floret and leaf tissue of most cultivars, whereas the broccoli stem showed an unusual pattern of accumulation. Sucrose exhibited greater cultivar dependency as well as seasonal variation compared to fructose and glucose in floret and leaf tissues. The floret tissue had a higher total content of sugar in the spring compared to the fall due to an increase in glucose and fructose. However, most of the leaf and stem tissues of broccoli had a higher total sugar content in the fall compared to the spring. Furthermore, stem and leaf tissues possessed a relatively higher total sugar content compared to floret tissue in both seasons. 'Grandeur' broccoli contained a significantly greater amount of total sugar in both floret and leaf tissues in both seasons, whereas 'YuDoRi No.1' broccoli exhibited the highest total content of sugar in stem tissue. At overall, the results showed significant influences of genotype, plant part and growing season on sugar content in broccoli.

Linking Leaf Functional Traits with Plant Resource Utilization Strategy in an Evergreen Scrub Species Rhododendron caucasicum Pall. along Longitudinal Gradient in Georgia (The South Caucasus)

  • Ekhvaia, Jana;Bakhia, Arsena;Asanidze, Zezva;Beltadze, Tornike;Abdaladze, Otar
    • Journal of Forest and Environmental Science
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    • v.38 no.2
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    • pp.110-121
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    • 2022
  • Leaf functional traits widely have been used to understand the environmental controls of resource utilization strategy of plants along the environmental gradients. By using key leaf functional traits, we quantified the relationships between leaf traits and local climate throughout the distributional range of Rhododendron caucasicum Pall. in eastern and western Georgian mountains (the South Caucasus). Our results revealed, that all traits showed high levels of intraspecific variability across study locations and confirmed a strong phenotypic differentiation of leaf functional variation along the east-west longitudinal gradient in response to the local climate; out of the explored climatic variables, the moisture factors related to precipitation and number of precipitation and dry days for winter and growth seasons were more strongly related to leaf trait variation than the elevation and air temperature. Among studied leaf traits, the leaf specific area (SLA) showed the highest level of variability indicating the different resource utilization strategies of eastern and western-central Rh. caucasicum individuals. High SLA leaves for western-central Caucasian individuals work in relatively resource-rich environments (more humid in terms of precipitation amount and the number of precipitation days in winter) and could be explained by preferential allocation to photosynthesis and growth, while eastern Caucasian samples work in resource-poor environments (less humid in terms of precipitation amount and the number of precipitation days in winter) and the retention of captured resources is a higher priority appearing in a low SLA leaves. However, more evidence from a broader study of the species throughout its distribution range by including additional environmental factors and molecular markers are needed for firmer conclusions of intraspecific variability of Rh. caucasicum.

Biosystematic Study on the Genus Viola in Korea - Comparative Morphology of the Viola albida Complex - (한국산 제비꽃속의 종분화론적 연구 - 태백제비꽃군의 비교형태학적 연구 -)

  • 김경식
    • Journal of Plant Biology
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    • v.34 no.3
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    • pp.229-238
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    • 1991
  • The main objective of this study was to clarify the taxonomic status and the speciation mechanism of Viola albida complex in Korea. The complex includes three very closely related taxa, taxonomic ranks of which are much in dispute. These taxa in the complex are sympatric in distribution, and they are very similar in most morphological characteristics. Comparative morphological analyses revealed that they can be distinguished on the basis of leaf shape, although they show complicated patterns of variation in leaf shape. The variation of leaf shape in each taxon appeared to be maintained through developmental process, and extensive cleistogamy and vegetative reproduction in these taxa are suggested as the mechanisms maintaining the variation. Genetic variation in this complex was monomorphic in all 14 loci examined. Palynological study also showed that these taxa have the same pollen morphology. These results strongly suggested that they should be recognized as varieties of V. albida; var, albida, var. chaerophylloides, and var. takahashii. The results also suggested that some individuals that can be assigned to var. takahashii might be hybrids between var. albida and var. chaerophylloides.

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Variation of the Morphological Characteristics in the Accessions of Zoysia Species and Their Hybrid Lines (Zoysiagrass 수집계통들과 종간교배계통들의 형태적 특성들의 변이)

  • 김형기;김기선;주영규;홍규현;김경남;이재필;모숙연;김두환
    • Asian Journal of Turfgrass Science
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    • v.10 no.1
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    • pp.1-11
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    • 1996
  • Fifty accessions in Zoysia species and forty-one hybrid lines were evaluated in order to detect analyze the genectic variation of Zoysiagrasses and to obtain the basic information for breeding. 1.Plant height ranged from 13 to 34.5cm, leaf length from 7 to 26cm, leaf width from 2 to 7mm, and first leaf height from 1.1 to 8.5cm 2. The correlations among plant height, leaf length and first-leaf height were high, but leaf Width was correlated only With leaf length. 3. Stolon length was varied from 10 to 108cm, number of stolons from 12 to 53, internode length from 2 to 6.1cm and coverage percentage from 10 to 90%. 4.There were correlation among stolen length, number of stolons, internode length and coverage percentage except between stolen and internode length. 5.M1J and MC2 showed the highest level in stolen length, number of stolons and coverage percentage. Therefore they will be the best breeding materials for the purpose of fast covering. 6.According to the analysis on October 26, 1995, yellow color-change rate of leaves ranged from 5 to 85%, red color-change from 0 to 75% and sum of yellow and red change from 20 to 95%. Speciallty M2M1 and FL-41 kept green color untill early December, indicating that they had the late dormancy. 7. Visual assessment of stolon length and number of stolons, showed similar tendency With actually measured data. 8. Shoot density was very high in M2M1 and S2, and showed negative correlation with leaf width. 9. It is known in Korea that the darker green color of lawngrass is, the better it looks. There were 11 lines with dark green color leaves such as J 85-10, Meyer in Z. japonira, one asacession in Z. matrella and M2M1, M2S8, JM2, 55C2 in hybrid lines.

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Characteristics of Leaf Morphology and Genetic Variation of the Rare Woody Plant, Berchemia racemosa var. magna (희귀수종 먹넌출 엽의 형태적 특성과 유전변이)

  • Song, Jeong-Ho;Lim, Hyo-In;Jang, Kyung-Hwan;Han, Jingyu
    • Korean Journal of Plant Resources
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    • v.26 no.5
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    • pp.613-618
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    • 2013
  • This study was conducted to investigate the morphological characteristics of leaf and the genetic diversity of Berchemia racemosa var. magna which is only found in Anmyeon Island of South Korea. ANOVA test showed that there were significant differences among individuals within population in all 10 leaf characteristics. Average characteristics of 39 individuals were 11.8 cm in leaf length, 7.1 cm in leaf width, 1.67 in leaf index, 5.4 cm in upper 1/3 width, 6.2 cm in lower 1/3 width, 3.6 cm in petiole length, 0.19 mm in leaf thickness, 11.5 ea. in number of veins (left), 11.4 ea. in number of veins (right) and 61.7 $cm^2$ in leaf area, respectively. Except for leaf thickness (18.8%), petiole length (21.7%) and leaf area (22.0%), the coefficients of variation of most leaf characteristics were relatively low (<15.0%). A total of 50 bands was generated from 8 selected I-SSR primers. The estimates of genetic variation were 1.719 in effective number of alleles ($A_e$), 26.0% in proportion of polymorphic bands (P), 0.410 in expected heterozygosity ($H_e$) and 0.598 in Shannon's diversity index (S.I.), respectively. In spite of the small number and the limited distribution, the B. racemosa var. magna population in Anmyeon Island showed high genetic diversity.

The Leaf Morphological Variation of Ten Regions of Natural Populations of Machilus thunbergii in Korea (후박나무 10개 천연집단의 엽형질 변이)

  • Yang, Byeong-Hoon;Song, Jeong-Ho;Lee, Jae-Cheon;Park, Young-Goo
    • Journal of agriculture & life science
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    • v.45 no.3
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    • pp.25-33
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    • 2011
  • This study was conducted to examine genetic variation on leaf characteristics of Machilus thunbergii populations. Ten populations were subjected to multivariate analysis for 9 characteristics of leaf morphology. Average length of leaf blade, leaf width, petiole length, vein number were 9.8cm, 4.0cm, 1.8cm, 8.4 respectively, while angle of leaf base and leaf apex were $67.9^{\circ}$ and $78^{\circ}$ respectively. The coefficient of variation (C.V.) on leaf characteristics was 20% which indicate similar features among the populations. Nested analysis showed statistically signigicant differences among populations as well as among individuals within populations. Genetic relationship between populations using complete linkage method showed four groups to Euclidean distance 1.2 and did not show a tendency to cluster into the same group. There were three principal components that had a meaningful eigenvalue over 1.0 among the 9 components. The explanatory power of the top three main components on the total variation was 92.8%. The first principal component (PC) was explained about 40.3% which is mainly correlated with maximum leaf width and the second PC was explained about 28.7% which is correlated with leaf blade length. The third PC was explained about 23.8% which is correlated with petiole length ($X_3$). These characters were important factors for analysis of the relationship among natural populations of M. thunbergii.

Seasonal Variation in Concentration and Composition of Monoterpenes from Artemisia princeps var. orientalis (쑥에 함유된 monoterpenes 의 함량과 조성의 계절적 변이)

  • Kim, Jong-Hee
    • The Korean Journal of Ecology
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    • v.19 no.4
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    • pp.321-328
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    • 1996
  • The profile and concentration of monoterpene metabolites in the leaf and stem of Artemisia princeps var. orientalis were quantified, and seasonal variation in monoterpenes of Artemisia plant was investigated. Samples were taken from five sites at the campus of Kyungnam University during maturing season. Monoterpenes in leaf and stem were analyzed using gas chromatography-mass spectrometry (GC-MS). The major constituents of A. princeps var. orientalis in both the leaf and stem were 21 monoterpenes.$ {\alpha}-pinene,\;{\beta}-pinene,\;{\beta}-myrcene$, dl-limonene, naphthalene and unknown monoterpenes with 5.49 and 16.27 of retenstion time were present in high concentrations of compounds identified on the leaf and stem of A. princeps var. orientalis. The cmounts of total monoterpenes of leaf were from two to five times higher than stem and rapidly decreased with the time, while that of stem was constnat except early spring. Most of the high percentage of monoterpenes in the leaf were those with later retention time. These results indicated that monoterpenes yields are considered to be more variable than monoterpene composition in responding to the time in both the leaf and stem.

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