• Horticultural Science & Technology
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• v.33 no.2
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• pp.155-165
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• 2015

Well-feathered (over 10 feathers) 'Fuji'/M.9 apple trees were planted at $3.0{\times}1.0m$ and trained to slender spindle with 2.5 m height or to tall spindle with 3.5 m height, and the vegetative growth, productivity, and fruit quality of two training systems were compared for 8 years. The canopy volume of the tall spindle trees surpassed that of the slender spindle trees 4 years after planting and was 25% larger than that of the slender spindle trees 5 years after planting. The accumulated yield over 8 years for the tall spindle system was 14% higher than that of the slender spindle system. Alternate bearing and incidence of marssonina blotch were observed in both treatments after 5 years of planting. There was often vegetative imbalance in the trees however, the degree of yield loss and vegetative imbalance of the tall spindle trees was lower than those of the slender spindle trees. Soluble solid content and fruit red color of the tall spindle trees were higher than that of the slender spindle trees in 5 year after planting, resulting from increased light penetration in the canopy due to even distribution of lateral branches and from fruit bearing in different height locations of the trees. In conclusion, increasing the tree height to about 3.5 m using slender spindle 'Fuji'/M.9 apple trees planted with over 333 trees per 10a led to better light penetration, yield and fruit quality compared to a conventional wide training system with the slender spindle.

• Journal of Bio-Environment Control
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• v.18 no.4
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• pp.492-501
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• 2009

This study was carried out to investigate influence of tree height on vegetable growth, productivity, and labor input in 5 year old slender spindle 'Fuji'/M.9 apple trees planted with the tree space of $3.2{\times}1.2m$ and trained to the tree height of 2.5m in the apple orchard of Apple Research Station, NIHHS, RDA in Gunwi County, Korea. Tree height was extended to the tree height of 4.0m for two years. And then, the tree height was restricted to 2.5 (control), 3.0, 3.5, and 4.0m in 8th year. Yield per 10a was 46, 25, and 4% higher at the tree height of 4.0, 3.5, and 3.0m than at the tree height of 2.5m respectively in first year and 17, 12, and 10% higher respectively, indicating the taller tree, the higher yield. Soluble solid contents was highest in the height of 2.5m than in any other treatments for 2 years, but fruit skin color was not clear. Labor input was increased parallel with the increase of tree height, but yield per labor input was decreased. Gross income and net income per 10a were increased parallel with the increase of tree height in first year, but not increased in second year. In considering production, labor input, and economic efficiency, the optimum tree height of slender spindle 'Fuji'/M.9 apple planted with the tree space of $3.2{\times}1.2m$ was presumed to be 3.0~3.5m.

• Journal of Bio-Environment Control
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• v.16 no.3
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• pp.258-263
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• 2007

This study was carried out to investigate the effects of number of the lower scaffold limbs on tree growth, light penetration, fruit yield, and fruit quality in slender spindle in 6-year-old 'Fuji'/M.9 apple trees. With regard to the growth by the numbers of the lower scaffold limbs, the width of the tree was wide and the growth of new shoots was increased when the number of the lower scaffold limbs was five. Compare with other treatments, five lower scaffold limbs showed high light-interception on the upper (150 cm above the ground) and middle (100 cm above the ground) canopy. There was no difference in the total number of the flower buds of the spurs according to the number of scaffold limbs, but the number and cross section area of flower bud on the lower canopy (120 cm above the ground) were increased where the number of the lower scaffold limbs was five. Fruit yield was highest in the treated with five lower scaffold limbs and fruit weight tended to increase where the number of the lower scaffold limbs was five or eight. With regard to fruit quality, there showed no difference in fruit shape index, firmness, acid content, Hunter L and b value according to the location of canopy and the number of the lower scaffold limbs, but the content of soluble solids was highest treated with five lower scaffold limbs. Hunter a value indicating fruit color was found to be highest treated with five lower scaffold limbs whose light interception was highest.

• Horticultural Science & Technology
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• v.32 no.4
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• pp.454-462
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• 2014

This study was carried out to determine the effect of tree height on light transmission, spray penetration, tree growth performance, fruit quality attributes, and labor productivity in the slender-spindle system of 'Hongro'/M.9 apple trees. With increasing tree height, the light penetration into the internal parts of the canopy decreased, especially in the lower canopy. Leaf area index (LAI) increased with increasing tree height, thereby leading to a reduction in the extent of spray penetration into the interior of the canopy. With increasing tree height, shoot growth was more vigorous but produced slender shoots in the upper canopy compared to the lower canopy. Although the soluble solid content and coloration of fruit decreased, there was no difference in fruit firmness and acidity. In addition, the number of final fruit set increased, although the production of large fruit (> 305 g) decreased. The increase in tree height also significantly increased the labor required for practices such as thinning of flowers and fruits, pruning, and harvesting. Nevertheless, this problem of increased in labor input in taller trees would was eased by use of a mechanical lift. Utilizing a lift for thinning the flowers of trees 4.5 m in height saved 14.6 min per tree, compared to the use of ladder. Therefore, it is highly considerable that in order to enhance light transmission and fruit coloration, light conditions should be improved in the internal tree canopy of slender-spindle systems.

• Horticultural Science & Technology
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• v.32 no.4
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• pp.440-447
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• 2014

For this analysis, canopies of slender-spindle 'Hongro'/M9 apple trees were divided into 4 positions: upper, lower, exterior, and interior parts. The photosynthetic rate was highest in the external part of the upper canopy of the tree, where it was 4.5 times higher than in the internal part of the lower canopy. This difference was closely associated with differences in light penetration based on canopy position. Analysis of leaf growth characteristics showed that the leaves situated in the internal part of the canopy were larger and thinner than those in the external part of the canopy. The difference in leaf thickness was mainly due to thickness of the primary layer of palisade tissues (68.5 and $110.3{\mu}m$ for internal and external leaves, respectively). Fruit weight and quality were closely related to the extent of light penetration. Fruit weight, soluble solid content, and red skin color were higher in the fruits from the external part of the canopy. Thus, fruit maturity was delayed in the internal part of canopy. The fruit skin and wax layer were thicker in fruits from the internal canopy than in those from the external canopy. Therefore, our results indicate a need for improved light penetration in internal parts of the canopy and for split harvesting depending on maturity at different canopy positions.

• Journal of Practical Agriculture & Fisheries Research
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• v.19 no.1
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• pp.39-49
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• 2017

The present study conducted a comparison on tree growth and productivity of high-density apple orchards by several orchard management systems: making scaffolds by heading-back and thinning out pruning (T-1), maintaining scaffolds upward and bearing shoots downward (T-2), managing branches slightly upward without heading-back (T-3), keeping leaders downward and shoots pending (T-4), maintaining leaders high and branches horizontal with severe pinching (T-5), making leaders with lower branches vigorous and upper shoots pending (T-6), and controlling very high planting density with bending branches (T-7). In conclusion, the orchards of (T-5) and (T-6) management systems showed a superior performance in controlling tree growth, productivity, and quality of fruits. Also, superior management efficiency was obtained in the orchards of (T-5) and (T-6).

• Applied Microscopy
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• v.29 no.4
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• pp.479-491
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• 1999

This study was performed to investigate the distribution and differentiation of choline acetyltransferase (ChAT)-immunoreactive cells in the basal nucleus of Meynert of the postnatal and adult rat forebrains, utilizing techniques of immunocytochemistry. According to the cell shape and the ratio of long axis vs short axis of cell soma, the ChAT-immunoreactive nerve cells in the basal nucleus of Meynert of the adult rat were classified into six types. In the adult rat, the frequency distributions (FD) of round, oval, elongated, fusiform, triangular and polygonal cells were 9.4%, 35.5%, 32.1%, 5.9%, 9.1% and 8.0%, respectively. The FD of oval and round nerve cells on the postnatal day (PND) 14 were observed to be 18.7% and 51.5%, respectively. Those were shown to be progressively decreased during developmental process to the adult. Also, those of elongated and triangular nerve cells on the PND 21 were observed to be 30.4% and 10.1%, respectively. Those were shown to be same phenomenon a,1 those in the round and oval cells. Meanwhile, those of the triangular and polygonal nerve cells were progressively increased from the early postnatal stage to the adult. The total mean volumes of ChAT-immunoreactive cell somata in the PND 7 rat were the lowest $(1,083{\mu}m^3)$ and those in the PND 21 rat were shown to be the highest $(5,045{\mu}m^3)$. But in the adult, those were decreased to $(2,731{\mu}m^3)$. Those in the PND 21 rat were shown to be about 84.7% larger than those in the adult. On the electron micrography, the cell organelles such as ribosomes, polysomes, rough endoplasmic reticula (RER) and mitochondria were well developed in the PND 21 rat forebrains, but Golgi complexes were shown to be proliferating phase. Especially, ribosomes, polysomes and RER were immunoreactive in the tissues treated with 0.05% triton X-100. According to the observations in the present study, it is considered that the ChAT-immunoreactive nerve cells in the basal nucleus of Meynert of the rat forebrains are differentiated throughout the following processes of changes during the postnatal development: 1) increase of cell soma volumes with the differentiation of tell organelles and neurites, 2) increase in the FD of differentiated tell types and 3) cell schrinkage without cell loss. The ribosomes, polysomes and RER are considered to be closely related to the intracellular localization and biosynthesis of the ChAT but not Colgi complex.

• Horticultural Science & Technology
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• v.28 no.3
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• pp.374-380
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• 2010

Well-feathered (5.2 feathers, stem diameter 13 mm) trees of 'Fuji' apple/ M.9 EMLA were planted at $4.0{\tiems}1.5$ m and whip trees (stem diameter 10 mm) of 'Fuji'/M.26 at $4.0{\times}2.0$ m were trained to the slender spindle. The productivity and yield efficiency of two orchard systems were compared for 6 years. The canopy volume of the tree/M.9 EMLA reached $2.07m^3$ in 2nd year and increased slowly to almost the targeted tree volume of $2.9m^3$ in 4th year. Trees/M.26 grew slowly at the begin but from 3rd year the tree volume expanded quickly to reach $5.6m^3$ in 5th year, covering over the allowed space. Yield of M.9 EMLA per 10a increased from 0.3 ton in 2nd year to 4.6 ton in 5th year, and 5.0 ton in 6th year, but yield of M.26 per 10a increased from 0.5 ton in 3rd to 2.9 ton in 6th year. Cumulative yield per 10a up to 6th year was 13.9 ton for M.9 EMLA but only 9.8 ton for M.26. Fruit weight for M.9 EMLA was heavier than that for M.26. In conclusion, the high density planting system with well-feathered trees/M.9 EMLA was better than the conventional wide planting system with whip trees/M.26.

• Horticultural Science & Technology
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• v.33 no.1
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• pp.1-10
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• 2015

This study was conducted in three years (7-9 years after planting) to investigate vegetative growth, yield, fruit quality, and return bloom for optimum crop load based on different planting densities of adult 'Fuji'/M.9 apple trees. As plant materials, 'Fuji'/M.9 apple trees planted at $3.5{\times}1.5m$ (190 trees per 10 a), $3.5{\times}1.2m$ (238 trees per 10 a), and $3.2{\times}1.2m$ (260 trees per 10 a) spacing and trained as slender spindles were used. The crop load was assigned to five different object ranges as follows: 55-64, 65-74, 75-84, 85-94, and 95-104 fruit per tree. TCA increment, total shoot growth, return bloom, yield per tree, and yield efficiency tended to increase as planting density decreased, and fruit weight and soluble solid content tended to increase as the object range of crop load decreased. Fruit red color tended to increase as shoot growth decreased. For apple trees planted with 238 trees and 260 trees per 10a, biennial bearing occurred when the crop load was over 85-94 and 75-84 fruits, respectively. However, biennial bearing did not occur when the crop load was 95-104 fruits in apple trees planted with 190 trees per 10a. Accumulated yield tended to increase as planting density and crop load increased, but that of biennial bearing did not show such a difference. Based on our results, optimum crop load recommendations are to set 95-104 fruits per tree in 'Fuji'/M.9 mature apple trees planted at 190 trees per 10a, 75-84 fruits per tree at 238 trees per 10a, and 65-74 fruits per tree at 260 trees per 10a.