• Title/Summary/Keyword: stem density

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Effect of Cell Size on Growth and Development of Plug Seedlings of Three Indigenous Medicinal Plants (플러그 셀 크기가 세 가지 자생 약용식물 묘 생육에 미치는 영향)

  • Oh, Hye Jin;Park, Yoo Gyeong;Park, Ji Eun;Jeong, Byoung Ryong
    • Journal of Bio-Environment Control
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    • v.23 no.2
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    • pp.71-76
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    • 2014
  • There have not been many studies conducted on the seedling production, especially in plug trays, of traditional medicinal plant species. In an effort to establish guide lines for seedling production, this study investigated the effect of plug cell size on the growth and development of plug seedling of three medicinal plant species. Seeds were sown in either 128, 200, or 288-cell plug trays, containing a commercial medium. Growth and development of individual seedling was generally promoted with increasing size of a plug cell in all of the three species. The greatest biomass of the seedlings gained in a plug tray was obtained in the 288-cell trays in Perilla frutescens var. acuta Kudo and Sophora tonkinensis, and the 200-cell trays in Angelica gigas Nakai. Overall growth and development of the shoot and root of a single seedling of Perilla frutescens var. acuta Kudo, except total chlorophyll and anthocyanin contents, was the greatest in the 128-cell tray. However, length of the longest root, length, width and area of the leaf, internode length, root fresh weight, and root ball formation in the 200- and 288-cell trays were not significantly different each other. In Sophora tonkinensis, although length of the longest root, stem diameter, leaf width, leaf area, shoot fresh weight, and root ball formation were not significantly different among the treatments, length of the longest root and root ball formation of a single seedling were the greatest in the 128-cell tray. Overall shoot and root growth, except total chlorophyll content, of a single seedling of Angelica gigas Nakai was the greatest in the 128-cell tray. Based on the total biomass, it is concluded that 288-cell trays are recommended for production of plug seedlings of medicinal plant species P. frutescens var. acuta Kudo and S. tonkinensis. In A. gigas Nakai, it would be more economical to use the 200-cell trays than 128-cell trays due to total biomass.

Changes in Growth Rate and Carbon Sequestration by Age of Landscape Trees (조경수목의 수령에 따른 생장율과 탄소흡수량 변화)

  • Jo, Hyun-Kil;Park, Hye-Mi
    • Journal of the Korean Institute of Landscape Architecture
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    • v.45 no.5
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    • pp.97-104
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    • 2017
  • Greenspace enlargement through proper landscape planting is essential to creating a low carbon society. This study analyzed changes in stem diameter growth rates(DGR), ratios of below ground/above ground biomass(B/A), and carbon sequestration by age of major landscape tree species. Landscape trees for study were 11 species and 112 individuals planted in middle region of Korea. The DGR and B/A were analyzed based on data measured through a direct harvesting method including root digging. The carbon sequestration by tree age was estimated applying the derived regression models. The annual DGR at breast height of trees over 30 years averaged 0.72 cm/yr for deciduous species and 0.83 cm/yr for evergreen species. The B/A of the trees over 30 years averaged 0.23 for evergreen species and 0.40 for deciduous species, about 1.7 times higher than evergreen species. The B/A by age in this study did not correspond to the existing result that it decreased as tree ages became older. Of the study tree species, cumulative carbon sequestration over 25 years was greatest with Zelkova serrata(198.3 kg), followed by Prunus yedoensis(121.7 kg), Pinus koraiensis(117.5 kg), and Pinus densiflora (77.4 kg) in that order. The cumulative carbon sequestration by Z. serrata offset about 5% of carbon emissions per capita from household electricity use for the same period. The growth rates and carbon sequestration for landscape trees were much greater than those for forest trees even for the same species. Based on these results, landscape planting and management strategies were explored to improve carbon sequestration, including tree species selection, planting density, and growth ground improvement. This study breaks new ground in discovering changes in growth and carbon sequestration by age of landscape trees and is expected to be useful in establishing urban greenspaces towards a low carbon society.

Effect of rice seed dressing with imidacloprid WS on early occurring rice insect pests (본답 초기해충의 생력방제를 위한 imidacloprid의 종자분의 효과)

  • Choi, Byung-Ryul;Yoo, Jae-Ki;Lee, Sang-Guei;Lee, Jeong-Oon
    • The Korean Journal of Pesticide Science
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    • v.2 no.1
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    • pp.85-90
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    • 1998
  • A series of experiments were carried out to determine the effect of seed dressing application of chemicals on the early occurring rice insect pests. The number of small brown planthopper adults per 3hills in seed-dressing plot of Imidacloprid WS (Im WS) was 1.6 and its control efficacy was over 90%, compared with that of untreated plot. Rice water weevil was observed in the density of 8.8 larvae per 5 hills seed-dressed Im WS plots(showing 95.5% of control efficacy). Control efficacy of Im WS and Im GR against adult weevils lasted for 26 days after treatment and was higher than that of Carbofuran GR. Rates of injured stems by rice stem maggot and injured leaves by rice leaf miner were 3% and 3.7% in Im WS treatment plot respectively. Seed germination rate after seed dressing with recommended dosage (3 g/seed kg) of Im WS was 71 % on the 1st day and increased on 5th day up to that of untreated seed. The residual amount of Im in seed dressing plot was 0.11 ppm in rice roots and 0.05 ppm in leaves on the 40th day after treatment. Residual effect of Im WSI sustained for 50 days with over 95% insecticidal effect for the rice water weevil and over 90% for the brown planthopper. Quantity of the chemical applied in the field was calculated as 0.084 kg a.i./ha in seed dressing and 0.3 kg a.i./ha in seed box treatment, respectively It took 1 hour to treat insecticide by seed dressing, 2.5 hours by seed box applying, and 3.6 hours by water surface releasing per 990 $m^2$.

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Studies on the Agricultural Use of the Water-swelling Polymer -I. Basic Experiment (수팽윤성(水膨潤性) 고분자(高分子) 화합물(化合物)의 농업적(農業的) 이용(利用)에 관(關)한 연구(硏究) -제(第) I 보(報) 기초시험(基礎試驗)을 중심(中心)으로)

  • No, Yeong-Pal;Jung, Yeun-Tae;Chung, Gun-Sik;Kim, Young-Ha
    • Korean Journal of Soil Science and Fertilizer
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    • v.20 no.3
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    • pp.209-216
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    • 1987
  • The experiments were carried out in lab. as well as in pots, to develop the agricultural usage of water swellable polymer, a kind of polyacrylic acid(K-sorb) synthesized by the Korea Advanced Institute of Science & Technology (KAIST) recently. The changes of soil physical properties and the influences to crops were investigated with various levels of K-sorb. When the K-sorb mixed with soils and soaked up distilled water, the volume of soils increased with the increase of soil available water contents and increase of K-sorb application levels. The rate increase of soil available water was higher in the coarse textured soils than in the fine while the swelling rate of soil volume showed adverse tendencies. A positive linear regression was observed between the contents of available soil water and levels of K-sorb. K-sorb application decreased bulk density and hardness due to the increase of porosity after soybean cultivation. The permeability in coarser textured soils such as sandy and coarse loamy families was decreased with the increase of K-sorb but in the medium textured soils it was opposite. At higher levels of K-sorb, about 0.5%, the permeability abruptly decreased due to dispersion and vertical movement in silty soils, while it was not changed in fine clayey soils but has the same trend with silty soils. In the plot of 0.3% of K-sorb application, the growth of soybean such as number of pods and stem length etc. increased and the yield also increased about 1.2-1.8 times of control. The optimum amounts of K-sorb were slightly different according to soil texture but estimated from regression curves were about 0.2% to 0.35% of soils in dry weight bases.

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Vegetation Change of Long-Term Ecological Research (LTER) Site of Abies koreana Forests in Hallasan Mountain (Yeongsil Area) (한라산(영실) 장기생태연구(LTER) 조사지 구상나무 숲 식생 변화)

  • Song, Kuk-Man;Kim, Jae-Hoon
    • Journal of the Korean Society of Environmental Restoration Technology
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    • v.21 no.1
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    • pp.1-12
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    • 2018
  • This study sought to analyze changes in the vegetation structure and species diversity indices in the Abies koreana forest in the Yeongsil area of Hallasan Mountain and establish basic data for Long-Term Ecological Research (LTER). The results show that the number of woody plants per 1ha has increased by 804 (45.1%) from 1,781 to 2,585 individual plants, whereas the number of live A. koreana did not change much with 796 individual plants. The number of dead tree plants, however, jumped by 807 (82.2%) from 982 to 1,789 plants. With other tree species, the number of live individual plants increased by 807 (82.2%) from 982 to 1,789 plants, whereas the number of dead plants increased by 31 from 10 to 41 individual plants. Results of the average importance index analysis show that the number of A. koreana decreased, while the number of Weigela subsessilis, Rhododendron yedoense for. poukhanense, and Magnolia sieboldii increased. The species diversity index increased by 6.7% as per the maximum level in the upper layer and by 7.2% in the lower layer, respectively. Results of the stem morphology analysis show that the number of AS type was highest with 700 individual plants among A. koreana, while their distribution by type shifted from AS>AL>DS>DB to AS>DS>AL>DB. As for other species, the AS type was most abundant with 1,580 individual plants, while their distribution by type shifted from AL>AS>AF>AB to AS>AL>AB>AD. The number of newly sprouted tree plants has increased from 420 in 2014 to 453 in 2016. Of the individual plants that were alive in 2014, 23 were found dead and 33 individual plants younger than 3 years old have been newly added to the group. The number of deaths after 2014 was highest among the 2 years olds, whereas the number of 2 years olds that sprouted in 2014 was highest among the newly sprouted tree plant group. In the A. koreana forest in Yeongsil, the density of other species was rising rapidly with the exception of the dead and live A. koreana. The occurrence of new tree plants is decreasing rapidly, thereby confirming the rapid change in the area's vegetation structure. Therefore, there is a need for continuous long-term ecological research to monitor vegetation change. It can also be used as a basic data for efficient restoration research.

The Variation of Natural Population of Pinus densiflora S. et Z. in Korea (IX) -Needle and Wood Characteristics of Six Populations- (소나무천연집단(天然集團)의 변이(變異)에 관(關)한 연구(硏究)(IX) -광주(廣州), 제천(堤川), 보은(報恩), 무주(茂朱), 구례(求禮), 제주집단(濟州集團)의 침엽(針葉) 및 재질형질(材質形質)-)

  • Yim, Kyong Bin;Lee, Kyong Jae
    • Journal of Korean Society of Forest Science
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    • v.44 no.1
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    • pp.1-25
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    • 1979
  • Six natural populations of Pinus densiflora S. et Z. as shown in the location map (Flg 1) were studied during 1978. The numerial pouplation codes, 13 to 18. The results of populations 1 to 12 were reported in previous papers. Following the study methods described before, 20 trees were sampled from each population and morpological characteristics such as stem forms, branching habit needle and wood properties investigated. The results are summerized as follows; 1. The mean stand ages were ranged from 36 to 97 of years. The growth performances of trees of population 14. 15 and 18 was similar, but 13, 16, and 17 seemed to be inferior more or less. 2. The ratios of clear bole length were 0.70 in population 18 as the highest but 0.28 for population 16 as the lowest. 3. The population 17 was considered to be a stand of the coarser branching habit having the crown index (The maximum crown diameter/the crown height) 158 though the branching angles were almost horizontal. 4. The differences were observed in the clear bole length ratios and crown-indices between population as shown In Fig. 3 and 4. 5. As to the serration density, number of stomata row and resin duct; the significant differences exist between individual trees within population and also between populations. 6. Population 18 shown resin duct index 0.119 as the maximum. 7. The patterns of diameter growth, based on the width of 10-year-ring segment unit(for example, the 1st segment denotes the with between pith center and 10th year ring and the 2nd one is from 11th to 20th ring and so on.), were alike among populations as shown in Fig 9. 8. Significant differences between population in mean summer wood percentage as well as in wood specific gravity was observed. The values of wood specific gravity were increased with the increase of ages in population 14, 18 however vice versa in population 13, 15, and 17. 9. The fiber length was mereased with the increased of age but no differences between populations as shown in Fig. 12.

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Effect of Shading Methods on Growth and Fruit Quality of Paprika in Summer Season (파프리카 여름재배시 차광방법이 생육과 과실특성에 미치는 영향)

  • Ha, Jun Bong;Lim, Chae Shin;Kang, Hyo Yong;Kang, Yang Su;Hwang, Seung Jae;Mun, Hyung Su;An, Chul Geon
    • Journal of Bio-Environment Control
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    • v.21 no.4
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    • pp.419-427
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    • 2012
  • This study was carried out to investigate the effect of two shading methods, shading agent spray on the glasshouse and internal shading screen treatment, on the growth and fruit quality of paprika (Capsicum annuum L. 'Cupra' and 'Coletti') in summer season cultivation. In the shading agent treatment, a commercial shading agent diluted with water at a ratio of 1 : 4 was sprayed on the roof of a glasshouse. In the internal shading screen treatment, a 10~20% shaded screen was used during the day time when the sun radiation was greater than $700W{\cdot}m^{-2}$. Compared to the unshaded control, photosynthetic photon flux density (PPFD) decreased in the greenhouse in the shading agent (SA) and shading screen (SS) treatments by 20% and 30%, respectively. Lower air temperatures and higher relative humidities were observed in the SA than in both the control and the SS treatment. Time to reach the break point of humidity deficit $8g{\cdot}m^{-3}$ was 2 hours late in the SA than in both the control and the SS treatment. Compared to control, both the SA and the SS treatments showed lower instantaneous temperatures of leaf, fruit, and flower by $2^{\circ}C$, $5^{\circ}C$ and $3^{\circ}C$, respectively. There were no differences in number of branches, stem diameter, and leaf size among treatments although both shading treatments promoted plant height in both cultivars. Botrytis infection ratio declined with the SA treatment by 14.7% in 'Cupra' and 22.1% in 'Coletti' as compared to that in the control. Shading increased fruit size in both cultivars, whereas no differences were observed in the number of locules and thickness of fruit tissue among treatments. Shading treatment increased mean fruit weight by a range of 10 to 15 g per fruit, while it decreased soluble solids contents as compared to that in the control. Similar Hunter values were observed among treatments, while fruit firmness increased slightly in shading treatments. Compared to the control, shading treatments improved marketable fruits by 11.7~22.6% and increased the number of fruits per plant by 4~9.2 in both 'Cupra' and 'Coletti'. The results of this study indicate that shading agent application on the roof of glasshouse would be one of the most effective options to reduce heat stress imposed on the paprika crop in summer cultivation, resulting in improved crop growth and fruit yield.

Measurement of Backscattering Coefficients of Rice Canopy Using a Ground Polarimetric Scatterometer System (지상관측 레이다 산란계를 이용한 벼 군락의 후방산란계수 측정)

  • Hong, Jin-Young;Kim, Yi-Hyun;Oh, Yi-Sok;Hong, Suk-Young
    • Korean Journal of Remote Sensing
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    • v.23 no.2
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    • pp.145-152
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    • 2007
  • The polarimetric backscattering coefficients of a wet-land rice field which is an experimental plot belong to National Institute of Agricultural Science and Technology in Suwon are measured using ground-based polarimetric scatterometers at 1.8 and 5.3 GHz throughout a growth year from transplanting period to harvest period (May to October in 2006). The polarimetric scatterometers consist of a vector network analyzer with time-gating function and polarimetric antenna set, and are well calibrated to get VV-, HV-, VH-, HH-polarized backscattering coefficients from the measurements, based on single target calibration technique using a trihedral corner reflector. The polarimetric backscattering coefficients are measured at $30^{\circ},\;40^{\circ},\;50^{\circ}\;and\;60^{\circ}$ with 30 independent samples for each incidence angle at each frequency. In the measurement periods the ground truth data including fresh and dry biomass, plant height, stem density, leaf area, specific leaf area, and moisture contents are also collected for each measurement. The temporal variations of the measured backscattering coefficients as well as the measured plant height, LAI (leaf area index) and biomass are analyzed. Then, the measured polarimetric backscattering coefficients are compared with the rice growth parameters. The measured plant height increases monotonically while the measured LAI increases only till the ripening period and decreases after the ripening period. The measured backscattering coefficientsare fitted with polynomial expressions as functions of growth age, plant LAI and plant height for each polarization, frequency, and incidence angle. As the incidence angle is bigger, correlations of L band signature to the rice growth was higher than that of C band signatures. It is found that the HH-polarized backscattering coefficients are more sensitive than the VV-polarized backscattering coefficients to growth age and other input parameters. It is necessary to divide the data according to the growth period which shows the qualitative changes of growth such as panicale initiation, flowering or heading to derive functions to estimate rice growth.

Effect of Shading and Supplemental Lighting for Greenhouse Cultivation of Cucumber in Summer Season (하절기 오이 온실재배 시 차광 및 보광 효과)

  • Jin Yu;Ji Hye Yun;So Yeong Hwang;Eun Won Park;Jeong Hun Hwang;Hyeong Eun Choi;Jeong Kil Koo;Hee Sung Hwang;Seung Jae Hwang
    • Journal of Bio-Environment Control
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    • v.32 no.3
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    • pp.226-233
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    • 2023
  • High solar radiation in summer season causes excessive respiration of crops and reduces photosynthesis. In addition, the rainy season, which mainly occurs in summer, causes a low light condition inside the greenhouse. A low light condition can reduce crop growth and yield. This study was conducted to evaluate the effect of shade and supplemental lighting on the growth and yield of cucumber during summer season. Cucumber grafted seedlings were transplanted in two plastic greenhouses on August 30, 2022. To reduce the light intensity inside the greenhouse, a 50% shading screen was installed in one greenhouse. Supplemental lighting was conducted from September 7, 2022 to October 20, 2022. HPS (high-pressure sodium lamp), W LED (white LED, red:green:blue = 5:3:2), and RB LED (combined red and blue LED, red:blue = 7:3) were used for supplemental lighting sources, and non-treated (nonsupplemental lighting) was as the control. The supplemental lighting was conducted before sunrise and after sunset for 2 hours with a photosynthetic photon flux density of 150 ± 20 µmol·m-2·s-1. The plant height, leaf length, leaf width, and SPAD value tended to increase in the shading group. RB LED increased stem diameter regardless of shading treatment. Fresh and dry weights of fruits were not significantly different in shading and supplemental lighting. Average fresh weight of fruits was not significantly different among supplemental lighting as the harvest date passed. In conclusion, in this study 50% shade treatment significantly improved the growth of cucumber during the summer season. In addition, the growth and fruit characteristics are better than the control without supplemental lighting. This study can be used as basic research data for applying supplemental lighting technology to cucumber cultivation.

The Variation of Natural Population of Pinus densiflora S. et Z. in Korea (III) -Genetic Variation of the Progeny Originated from Mt. Chu-wang, An-Myon Island and Mt. O-Dae Populations- (소나무 천연집단(天然集團)의 변이(變異)에 관(關)한 연구(硏究)(III) -주왕산(周王山), 안면도(安眠島), 오대산(五臺山) 소나무집단(集團)의 차대(次代)의 유전변이(遺傳變異)-)

  • Yim, Kyong Bin;Kwon, Ki Won
    • Journal of Korean Society of Forest Science
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    • v.32 no.1
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    • pp.36-63
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    • 1976
  • The purpose of this study is to elucidate the genetic variation of the natural forest of Pinus densiflora. Three natural populations of the species, which are considered to be superior quality phenotypically, were selected. The locations and conditions of the populations are shown in table 1 and 2. The morphological traits of tree and needle and some other characteristics were presented already in our first report of this series in which population and family differences according to observed characteristics were statistically analyzed. Twenty trees were sampled from each populations, i.e., 60 trees in total. During the autumn of 1974, matured cones were collected from each tree and open-pollinated seeds were extracted in laboratory. Immediately after cone collection, in closed condition, the morphological characteristics were measured. Seed and seed-wing dimensions were also studied. In the spring of 1975, the seeds were sown in the experimental tree nursery located in Suweon. And in the April of 1976, the 1-0 seedlings were transplanted according to the predetermined experimental design, randomized block design with three replications. Because of cone setting condition. the number of family from which progenies were raised by populations were not equal. The numbers of family were 20 in population 1. 18 in population 2 and 15 in population 3. Then, each randomized block contained seedlings of 53 families from 3 populations. The present paper is mainly concerned with the variation of some characteristics of cone, seed, needle, growth performance of seedlings, and chlorophyll and monoterpene compositions of needles. The results obtained are summerized as follows. 1. The meteorological data obtained by averaging the records of 30 year period, observed from the nearest station to each location of populations, are shown in Fig. 3, 4, and 5. The distributional pattern of monthly precipitation are quite similar among locations. However, the precipitation density on population 2, Seosan area, during growing season is lower as compared to the other two populations. Population 1. Cheong-song area, and population 3, Pyong-chang area, are located in inland, but population 2 in the western seacoast. The differences on the average monthly air temperatures and the average monthly lowest temperatures among populations can hardly be found. 2. Available information on the each mother trees (families) studied, such as age, stem height, diameter at breast height, clear-bole-length, crown conditions and others are shown in table 6,7, and 8. 3. The measurements of fresh cone weight, length and the widest diameter of cone are given in Tab]e 9. All these traits arc concerned with the highly significant population differences and family differences within population. And the population difference was also found in the cone-index, that is, length-diameter ratio. 4. Seed-wing length and seed-wing width showed the population differences, and the family differences were also found in both characteristics. Not discussed in this paper, however, seed-wing colours and their shapes indicate the specificity which is inherent to individual trees as shown in photo 3 on page 50. The colour and shape are fully the expression of genetic make up of mother tree. The little variations on these traits are resulted from this reason. The significant differences among populations and among families were found in those characteristics, such as 1000-seed weight, seed length, seed width, and seed thickness as shown in table 11. As to all these dimensions, the values arc always larger in population 1 which is younger in age than that of the other two. The population differences evaluated by cone, seed and seed-wing sizes could partly be attributed to the growth vigorousity. 5. The values of correlation between the characteristics of cone and seed are presented in table 12. As shown, the positive correlations between cone diameter and seed-wing width were calculated in all populations studied. The correlation between seed-wing length and seed length was significantly positive in population 1 and 3 but not in population 2, that is, the r-value is so small as 0.002. in the latter. The correlation between cone length and seed-wing length was highly significant in population 1, but not in population 2. 6. Differences among progenies in growth performances, such as 1-0 and 1-1 seedling height and root collar diameter were highly singificant among populations as well as families within population(Table 13.) 7. The heritability values in narrow sense of population characteristics were estimated on the basis of variance components. The values based on seedling height at each age stage of 1-1 and 1-0 ranged from 0.146 to 0.288 and the values of root collar diameter from 0.060 to 0.130. (Table 14). These heritability values varied according to characteristics and seedling ages. Here what must be stated is that, for calculation of heritability values, the variance values of population was divided by the variance value of environment (error) and family and population. The present authors want to add the heritability values based on family level in the coming report. It might be considered that if the tree age is increased in furture, the heritability value is supposed to be altered or lowered. Examining the heritability values studied previously by many authors, in pine group at age of 7 to 15, the values of height growth ranged from 0.2 to 0.4 in general. The values we obtained are further below than these. 8. The correlation between seedling growth and seed characteristics were examined and the values resulted are shown in table 16. Contrary to our hypothetical premise of positive correlation between 1-0 seedling height and seed weight, non-significance on it was found. However, 1-0 seedling height correlated positively with seed length. And significant correlations between 1-0 and 1-1 seedling height are calculated. 9. The numbers of stomata row calculated separately by abaxial and adaxial side showed highly significant differences among populations, but not in serration density. On serration density, the differences among families within population were highly significant. (Table 17) A fact must be noted is that the correlation between stomata row on abaxial side and adaxial side was highly significant in all populations. Non-significances of correlation coefficient between progenies and parents regarding to stomata row on abaxial side were shown in all populations studied.(Table 18). 10. The contents of chhlorophyll b of the needle were a little more than that of chlorophyll a irrespective of the populations examined. The differences of chlorophyll a, b and a plus b contents were highly significant but not among families within populations as shown in table 20. The contents of chlorophyll a and b are presented by individual trees of each populations in table 21. 11. The occurrence of monoterpene components was examined by gas liquid chromatography (Shimazu, GC-1C type) to evaluate the population difference. There are some papers reporting the chemical geography of pines basing upon monoterpene composition. The number of populations studied here is not enough to state this problem. The kinds of monoterpene observed in needle were ${\alpha}$-pinene, camphene, ${\beta}$-pinene, myrcene, limonene, ${\beta}$-phellandrene and terpinolene plus two unknowns. In analysis of monoterpene composition, the number of sample trees varied with population, I.e., 18 families for population 1, 15 for population 2 and 11 for population3. (Table 22, 23 and 24). The histograms(Fig. 6) of 7 components of monoterpene by population show noticeably higher percentages of ${\alpha}$-pinene irrespective of population and ${\beta}$-phellandrene in the next order. The minor Pinus densiflora monoterpene composition of camphene, myrcene, limonene and terpinolene made up less than 10 percent of the portion in general. The average coefficients of variation of ${\alpha}$-pinene and ${\beta}$-phellandrene were 11 percent. On the contrary to this, the average coefficients of variation of camphene, limonene and terpinolene varied from 20 to 30 percent. And the significant differences between populaiton were observed only in myrcene and ${\beta}$-phellandrene. (Table 25).

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