• Journal of Korean Society of Forest Science
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• v.98 no.3
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• pp.339-350
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• 2009

This study was carried out to investigate distributional condition of rare plants and useful plant resources, and to verify distributional patterns of understory vegetation associated with the upper layer's vegetation structure. Total 59 families, 160 genera, 218 kinds of vascular plants were identified at the study site including 6 rare plants designated by Korea Forest Service (Lloydia triflora Bak., Trillium kamtschaticum Pall., Lilium distichum Nakai, Anemone koraiensis Nakai, Iris odaesanensis Y.N. Lee, Viola diamantica Nakai). Twenty three species of useful plant resources were also identified at the site; 8 of them showed clustered distributions and the others were prone to scatter. Actual vegetation of this study area consisted of one natural community dominated by Quercus mongolica Fisch. and three disturbed communities of Larix kaempferi (Lamb.) Carriere, Abies holophylla Max. and/or a herbaceous vegetation resulting from forest removal and strong wind of mountain top. This classification was strongly supported by cluster analysis based on the surveyed plot data. Distributional patterns of understory vegetation within forest stand were somewhat related to overstory vegetation structure, but showed a different tendency according to site condition, species composition, and competitive pressure among understory vegetation. Therefore, in order to protect the important understory components as forest genetic resources, forest treatments such as density control of overstory should be implanted based on understanding of impact on understory's dynamics and growing condition.

• Journal of Ginseng Research
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• v.32 no.4
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• pp.347-356
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• 2008

Forest-grown American ginseng (Panax quinquefolius L.) is exposed to daily and seasonal light variations. Our goal was to determine the effect of understory light changes on the maximum quantum yield of photosystem II, expressed as $F_v/F_m$, and photosynthetic pigment composition of two-year-old plants. Understory light photon flux density and sunfleck durations were characterized using hemispherical canopy photography. Our results showed that understory light significantly affected the $F_v/F_m$ of American ginseng, especially during the initial development of the plants when light levels were the highest, averaging 28 mol $m^{-2}d^{-1}$. Associated with low $F_v/F_m$ during its initial development, American ginseng had the lowest levels of epoxidation state of the xanthophyll cycle of the season, suggesting an active dissipation of excess light energy absorbed by the chlorophyll pigments. As photon flux density decreased after the deployment of the forest canopy to less than 10 mol $m^{-2}d^{-1}$, chlorophyll a/b decreased suggesting a greater investment in light harvesting pigments to reaction centers in order to absorb the fleeting light energy.

• Journal of Korean Society of Forest Science
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• v.98 no.5
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• pp.563-567
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• 2009

This study was conducted to clarify the characteristics of bird communities between before and after forest fire in tropical deciduous Dipterocarp forest of Mondulkiri protected forest of the northeastern Cambodia from January to April 2009. The DBH distribution of trees were different in each DBH class. Most of the trees (> 80%) were belong to < 30 cm DBH. After the forest fire, coverage of understory layers were dramatically decreased by the fire. Total 64 species of birds were recorded, and 64 and 46 species of birds were observed before and after the fire, respectively. Observed number of individuals of bee-eaters, treepies, kingfishers, lapwings, herons, junglefowl, peafowl, prinias and warblers were decreased after the fire. The decrease of those species would be related with the change of habitat condition, such as decrease of water amount and understory coverage. For the management and conservation of junglefowls, peafowls, prinias and warblers, understory vegetation should be maintained in Mondulkiri protected forest, northeastern Cambodia.

• Journal of Ecology and Environment
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• v.43 no.2
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• pp.202-211
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• 2019

Oceanic islands are biologically important for their unique assemblages of species and high levels of endemism and are sensitive to environmental change because of their isolation and small species source pools. Habitat destruction caused by human landscape development is generally accepted as the main cause of extinction on islands, with exotic species invasion a secondary cause of extinction, especially on tropical islands. However, secondary impacts of human development (e.g., general degradation through resource use and exotic species introduction) are understudied on temperate islands. To determine secondary impacts of human development on the understory vegetation community, 90 field sites on Ulleung Island, South Korea, were sampled during the summer of 2016. Understory vegetation was chosen as it is a proxy for ecosystem health. Diversity and percent cover of introduced, native, and endemic species were tested against proximity to developed areas and trail usage using a model selection approach. Diversity was also tested against percent cover of three naturalized species commonly found in survey plots. The main finding was that distance to development, distance to town, and trail usage have limited negative impacts on the understory vegetation community within best-supported models predicting native and introduced cover and diversity. However, endemic species cover was significantly lower on high usage trails. While there are no apparent locally invasive plant species on the island at the time of this study, percent cover of Robinia pseudoacacia, a naturalized tree species, negatively correlated with plot diversity. These findings indicate that forests on Ulleung Island are not experiencing a noticeable invasion of understory vegetation, and conservation efforts can be best spent preventing future invasions.

• Journal of Korean Society of Forest Science
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• v.97 no.6
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• pp.597-604
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• 2008

Recognizing the necessity of the development of ecologically sound silvicultural system for the mature Korean pine (Pinus koraiensis), this study was carried out to examine the invasion and growth of understory vegetation after partial cutting. In 1997, the age class VII Korean pine forest in the Experimental Forest of Kangwon National University was opened up about 50% by modified shelterwood cutting with site preparation to induce succeeding regenerations. After 10 years, plenty of hardwood have come to the stand, forming understory vegetation. The abundance and composition of understory vegetation was periodically investigated in the ten $10m{\times}10m$ permanent sample plots, estimating importance values, species diversity index and the distribution of diameter and height of the understory vegetation. Encouraged by canopy opening, the coverage of understory reached perfect closure, composed of 22 tree species and 20 shrub species in 2007. Morus bombycis occupied 29.6% of the total importance value and Cornus controversa was 17.0%, followed by Styrax obassia, Quercus aliena, Quercus mongolica, Acer mono, and Pinus koraiensis. In the year of 2000 the species diversity was highest as 2.547 with 26 tree species and deceasing thereafter, showing 2.220 with 22 tree species in 2007. As understory layer was thickly covered and got grow bigger, some shade intolerant species were disappeared, lowering species diversity. In 2007 the biggest tree was a Quercus aliena with 11.3 cm of DBH and the tallest tree was a Cornus controversa with 9.8m of height. The frequency distribution of number of trees by diameter and height classes formed the inverted-J-shaped curve, supposed to be typical uneven-aged stand.

• Journal of Korean Society of Forest Science
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• v.100 no.1
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• pp.34-41
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• 2011

Vegetation structure, composition and diversity were quantified for 10 samples ($10m{\times}10m$) representing woody vegetation and for 30 samples ($1m {\times}3m$) representing understory vegetation in Pinus parviflora and Tsuga sieboldii forest of Taeharyeong, Ulleung-gun (Gyeongsangbuk-do). P. parviflora was limitedly advanced to sapling layer from seedling stage, and based on Mantel tests, composition of canopy layer was not established in ground woody vegetation. Non-metric multidimensional scaling revealed influence of biotic and abiotic factors in species composition of woody and understory vegetation. In the result of multiple regression model, abundance of P. parviflora (density and breast height area) and percent cover of woody debris were significant predict variables for understory diversity. These results suggest that relatively large disturbance is required for regeneration of P. parviflora and T. sieboldii forest, and control of expansion of monocultural understory species that monopolize resources such as Carex blepharicarpa and Maianthemum dilatatum, is necessary for maintenance of diversity.

• Animal cells and systems
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• v.5 no.4
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• pp.319-325
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• 2001

Transport of colored papers and decomposition of leaf litter of Quercus mongolica, Acer pseudo-sieboldianum, and Kalopanax pictus were investigated on three patches differentiated by aspect and understory in a temperate hardwood forest. Two patches are represented by dwarf bamboo (Patch SS) and herbaceous plants (Patch S), respectively, Iying on a south-west-facing slope. The other patch (Patch N) is located on a northeastfacing slope with herbaceous plants. Colored papers were placed on the patches to understand the pattern of litter movement on the ground. Papers were move dispersed in Patch S than in the other two patches. Some of the colored papers placed in Patch S moved upward. The results suggest that the litter movement is affected by aspect and that the leaf litter is retained by dwarf bamboo in Patch SS. Decay constant of Q. mongolica was significantly (p＜0.05) lower than those of K. pictus and A. pseudo-sieboldianum. Decay rates of Q. mongolica were significantly different between Patches N and S and between Patches SS and S (p＜0.05). On the other hand, decay rates of the other species were not significantly different among the three patches. The results suggest that aspect and understory exert an influence on redistribution and decomposition of leaf litter and that the effects could be different among the plant species.

• The Korean Journal of Ecology
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• v.11 no.2
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• pp.83-95
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• 1988

Phytomass, annual net production, respiration, annual gros production and litter fall were estimated in young Korean pine (Pinus koraiensis) plantations reforested 1,2,3,6,9 and 11 years ago, respectivly. The estimated phytomasses in terms of allometry were 0.07, 0.44, 0.58, 13.37, 32.26 and 48.42ton/ha/yr for annual net production and those of understory with harvest method were 3.70, 4.52, 5.28, 6.72, 7.56 and 5.72ton/ha for standing phytomass and 5.01, 4.71, 5.52m 6.15, 4.51 and 2.08 ton/ha/yr for annual net production at 1,2,3,6,9 and 11 year old plantation, respectively. Although the standing phytomass of understory increased gradually until 9 years thereafter decreased, that of white pine increased suddenly from 6 years after reforestation. The ratios of the annyal net production of understory to that of white pine were 71.57, 1.48 and 0.23 at 1,6 and 11 year old plantations, respectively. The net production in calorie value for whole vegetation of the plantations, respectively. The net production in calorie value for whole vegetation of the plantations increased until 9 years thereafter decreased; i.e. 224, 221, 280, 466, 527 and 494X105 kcal/ha/yr at 1,2,3,6,9 and 11 year old plantation, respectivey. The ratio of annual net production to annual gross production of the undersoty was lower than that of the pine at the same plantation age. Efficiency of solat energy untilization for net production by the pine was lower than that by understory y\until six ear old plantation but that was vice versa over 9 year old plantation. That for gross producrion by the pine was lower than that by the unsersotry at all plantations.

• Journal of the Korean Society of Environmental Restoration Technology
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• v.21 no.5
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• pp.81-96
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• 2018

The purpose of this study is to analyze the characteristics of vegetation changes and structures in the buffer green zone of Sihwa Industrial Complex and propose environmental, ecological and multi-functional vegetation management directions. The density of the Pinus thunbergii decreased from $23.1trees/100m^2$ in 2006 to $9.6trees/100m^2$ in 2017 due to the influence of thinning works. Its green volume increased from $0.97m^3/m^2$ to $2.02m^3/m^2$, contributing to the improvement of the atmospheric environment and promotion of biodiversity. The density of deciduous broad-leaved trees at the top maintained at $18{\sim}21trees/100m^2$. In the areas where the understory vegetation was removed, Pueraria lobata spread as an invasive plant and disturbed the forest. In the areas where the understory vegetation was not removed, various native plant species including the Rhus javanica and Celtis sinensis were introduced and contributed to the development of a forest in a multiple structure. There was a concern with the spread of Robinia pseudoacacia which developed into understory vegetation in Pinus thunbergii forest and some canopy forest. The study proposed vegetation management directions for the back, top and front side according to the physical structure of large-scale buffer green zone formed in a long linear form and the usage characteristics of adjacent land.

• Journal of the Korean Society of Environmental Restoration Technology
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• v.13 no.2
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• pp.80-94
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• 2010

The purpose of this study is suggest to restoration model of Pinus thunbergii in Saha-gu, Busan. The result of this study is summarized as follows. As the results of this study, vegetation restoration model is presented by separating community planting and edge planting. In community planting, as a group of canopy, there are 6 species; Pinus thunbergii, Quercus acutissima, Quercus dentata, Quercus serrata, Quercus alienna, Quercus variabilis. As a group of understory, there are 5 species; Platycarya strobilacea, Prunus sargentii, Styrax japonica, Eurya japonica, Morus bombycis. Also as a group of shrub, there were 15 kinds of species; Ulmus pavifolia, Ulmus davidiana, Lindera obtusiloba, Elaeagnus macrophylla, Mallotus japonicus, Ligustrum obtusifolium, Sorbus alnifolia, Rhus trichocarpa, Zanthoxylum schinifolium, Rosa wichuraiana, Rhus chinensis, Viburnum erosum, Rhododendron mucronulatum, Rhododendron yedoense, Indigofera pseudotinctoria. And as a group of edge vegetation, there were 10 kinds of species; Japanese Angelica, Symplocos chinensis, Pittosporum tobira, Lespedeza maximowiczii, Lespedeza bicolor, Rubus coreanus, Rubus idaeus, Vitis thunbergii, Ampelopsis brevipedunculata, Rosa multiflora. Vegetation restoration models of Pinus thunbergii community were calculated the units $400m^2$ for the average populations of the woody layer is 24 in canopy layer, 35 in understory layer, 410 in shrub layer, 34% herbaceous layer ground cover. And the average of breast-high area and canopy area is $10,852cm^2$ in canopy layer, in understory layer $1,546cm^2$, in shrub layer $1,158,660cm^2$. The shortest distance between trees is calculated as 2.0m in canopy layer, 1.9m in understory layer.