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Effects of Simulated Acid Rain on Nutrient Contents of Pinus densiflora S. et Z. and Forsythia koreana Nak. Seedlings (인공산성우(人工酸性雨)가 소나무 및 개나리묘(苗)의 식물체내(植物體內) 함유성분(含有成分)에 미치는 영향(影響))

  • Cheong, Yong Moon
    • Journal of Korean Society of Forest Science
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    • v.77 no.3
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    • pp.259-268
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    • 1988
  • For the purpose of examining the effects of simulated acid rain on nutrient contents of plant tissues in Pintos densiflora seedlings and Forsythia koreana rooted cuttings, the experimental design of randomized block arrangement with three replications was implemented in the experimental field of Yesan National Agricultural Junior College. One-year-old Pinus densiflora seedlings and Forsythia koreana cuttings were planted in the pots filled the mixed soils(nursery soil : forest soil of siliceous sandy loam=1 : 1 v/v) in the early spring of 1986. The regime of artificial acid rain, in terms of spray frequency per month and spray amount at single treatment per plot, was simulated on the basis of climatological data averaged from 30 years records. Simulated acid rain(pH 2.0, pH 4.0, and pH 5.5 as control) containing sulfuric and nutric acid in the ratio of 3 : 2(chemical equivalant basis) diluted with ground water were treated on the experimental plants under condition of cutting off the natural precipitation with vinyl tunnel, during the growing season(May 1 to August 31) in 1986. The results obtained in this study were as follow : (1) As for the nitrogen contents in plant tissues, P. densiflora increased significantly in one-year-old stembranch and root tissues, and F. koreana increased significantly in leaf and root tissues, as the pH levels of acid rain decreased. (2) The available phosphate contents in root tissues of P. densiflora, and in leaf and root tissues of F. koreana were significantly decreased, as the pH levels of acid rain decreased. (3) $K_2O$, CaO and MgO contents in plant tissues were significantly decreased in the both species as the pH levels decreased. And the effects of acid rain on F. koreana were higher than those of P. densiflora. (4) Sulfur contents of plant tissues in the both species were increased at pH 2.0 treatment. There were significant differences among three acid rain treatments in leaf and root tissues of P. densiflora, and in all parts of F. koreana. (5) In the effects of simulated acid rain on the both species and the tested soils, in general, F. koreana revealed higher sensitiveness than P. densiflora, and the lower pH levels of simulated acid rain were treated, the more sharp reaction was showed.

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Comparison of Thatch Accumulation in Warm-Season and Cool-Season Turfgrasses under USGA and Mono-layer Soil Systems (USGA 지반 및 약식지반에서 난지형과 한지형 잔디의 대취축적 비교)

  • Kim, Kyoung-Nam;Kim, Byoung-Jun
    • Journal of the Korean Institute of Landscape Architecture
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    • v.38 no.1
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    • pp.129-136
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    • 2010
  • This study was initiated to investigate thatch accumulation in several turfgrasses grown under two soil systems. The 45 centimeter deep USGA system was constructed with rootzone, intermediate and drainage layers. The mono-layer system, however, was made with only a 30cm rootzone layer. Turfgrasses used in the study were comprised of 3 varieties from Korean lawngrass of Warm-Season Grass(WSG) and 3 blends and 3 mixtures from Cool-Season Grass(CSG). A total of 9 turfgrass treatments were replicated three times in RCBD in both systems. Cultural practices for the research plot followed a typical maintenance program for highly managed turf. Treatment differences for thatch accumulation were observed among the turfgrasses in both soil systems. Thatch under the USGA system was 9% greater than under the mono-layer system due to its more favorable conditions for turf growth. Higher thatch depth was found with Korean lawngrass, 34~87% in the USGA system and 16~75% in the mono-layer system when compared with CSG. Among WSG, the Joongji variety was the highest in thatch layer under both the USGA and mono-layer systems. Kentucky bluegrass(KB) was the greatest among CSG, since it is a rhizomatous-type in growth habit, resulting in faster production of organic matter over bunch-type of tall fescue and perennial ryegrass. Proper depth in the thatch layer was known to be beneficial by enhancing the resiliency and wear tolerance of the turf in athletic fields. Thus, KB was considered to be a very excellent turfgrass in terms of turf quality, environmental performance, physical properties and soccer player safety. However, disadvantages such as poor water-holding properties, more inclined to injury from environmental stresses and severe diseases and insect injury were also expected where thatch was excessively accumulated. Therefore, these results demonstrate that more frequent measures for controlling thatch such as vertical mowing, topdressing or coring should be employed for soccer fields with Korean lawngrass and KB over other turfgrasses.

Plot Size for Investigating Forest Community Structure (V) - Adequate Number of Plots for Tree and Shrub Strata in a Mixed Forest Community of Broad-leaved Trees at Guryongsan Area - (산림군집구조 조사를 위한 조사구 크기에 관한 연구(V) - 구룡산지역 활엽수혼효림군집 교목층과 관목층의 적정 조사구수 -)

  • 박인협;서영권
    • Korean Journal of Environment and Ecology
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    • v.15 no.4
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    • pp.394-400
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    • 2002
  • A mixed forest community of broad-leaved trees in Guryongsan area was studied to determine the adequate number of plots of tree and shrub strata for investigating forest community structure. Fifteen 10m$\times$10m plots were set up in the tree stratum and fifteen 5m$\times$5m plots were set up in the shrub stratum. Species-area curves and performance curves were made from vegetation analysis of the plots. In tree stratum, the minimum number of plots where a given percentage increase in number of plots produced less than the same percentage in number of species was five. In tree stratum, minimum number of plots where a given percentage increase in nuts her of plots produced less than the half of the percentage increase in number of species was ten. In tree stratum, minimum number of plots where the dominant species was distinguished from the subdominant species was four In tree stratum. minimum number of plots where the subdominant species were distinguished from each other was eight. In tree stratum, the difference of species diversity(H') between seven or more plots and total fifteen plots was less than 0.05 Similarity index was more than 80% between aye or more plots and total fifteen plots, and more than 90% between nine or more plots and total fifteen plots. In shrub stratum, the minimum number of plots where a given percentage increase in number of plots produced less than the same percentage in number of species was five . In shrub stratum, the minimum number of plots where a given percentage increase in number of plots produced less than the half of the percentage increase in number of species was twelve. In shrub stratum. minimum number of plots where the dominant species was distinguished from the subdominant species was four. In shrub stratum, the minimum number of plots where the subdominant species were distinguished from each other was thirteen. In shrub stratum, the difference of species diversity(H') between six or more plots and total fifteen plots was less than 0.05. Similarity index was more than 80% between five or more plots and total fifteen plots, and more than 90% between nine or more plots and total fifteen plots. It may be given as a conclusion that adequate number of 10m$\times$10m plots for the tree stratnm was about 5 in general acse and 10 in case of requiring more accuracy, and that the adequate number of 5m$\times$5m plots for ahrub stratum was about 5 in general case and 13 in case of requiring more accuracy.

Effect of Shading Levels on the Soil Properties, Growth Characteristics, and Chlorophyll Contents of Ligularia stenocephala (차광정도가 곤달비의 토양변화, 생육상황 및 엽록소 함량에 미치는 영향)

  • Park, Byoung-Mo;Kim, Chang-Hwan;Bae, Jong-Hyang;Shin, Jung-Ryeul
    • Journal of Bio-Environment Control
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    • v.20 no.4
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    • pp.352-356
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    • 2011
  • It is true that the industrial development has usually been accompanied with urbanization or centralization of population that has inevitably led to high-rise buildings and densely built-up living area in the cities. While it is badly needed to acquire as much green land within the city limits as possible to compensate for reduced space for recreational purpose in parallel with increasingly urbanized area, the living conditions of plants have become seriously devastated due to shortage of sun light walled-off by high-rise buildings and contaminated environment and air. The shade that is generated by high-rise and compact buildings hinders growth of plants, which makes it urgent to develop native ground cover plant that is strongly viable in the shade. For this purpose, Ligularia stenocephala, best known as greens for Ssam (rice and condiments wrapped in leaves) was cultivated under the 30%, 50%, and 80% shadings and observed to see if there would be any changes in soil conditions, growth of plants and chlorophyll contents depending on the shading rate. The leaf number was 10.8 pieces under the 50% shading and 8.4 under the 30%-shading, 7.7 pieces more than that cultivated under lighting. The leaf width turned out to be excellent from cultivation under the 50%- shading, an evidence indicating its possibility of being cultivated as native ground cover plant in the shade. The live weight of the plants cultivated under the shading increased to 31.63 g, 43.39 g and 19.40 g, respectively, compared to 90.43 g of those in the untreated control plot. The increase in growth of roots was particularly significant with 48.48 g in comparison to 12.33 g under 30% shading cultivation. The chlorophyll synthesis amounted to 46.2 under the 50% shading, showing an increase compared to 41.9 under lighting. The chlorophyll synthesis rather shrank under other shading conditions. The cultivation of Ligularia stenocephala under the 50% shading showed the best condition in growth as native ground cover plant.

STUDIES ON THE DIMORPHISM AND TRANSITION OF BISEXUALITY OF HETEROSTYLOUS POLYGONACEAE (여뀌과 이형경식물의 Dinorphism과 Bisexuality의 변화)

  • Harn, Chang-Yawl
    • Journal of Plant Biology
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    • v.3 no.2
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    • pp.6-18
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    • 1960
  • The present experiments were designed in order to clarify the differences between the long and short styled plants and the transgressive gradition in the degree of dimorphism among the three heterostylous species of the Polygonus, P. japonica, F. esculentum, and P. senticosa, based on investigations regarding the floral structure, ecological and physiological traits, the results of which are summarized as follows: (1) P. japonica, although it exhibits typical dimorphism, has undergone so high a differentiation between long and short styled that its long styled individuals behave as if they were female; and short styled individuals as if male. In long-styled individuals, filament, anther, and pollen grains show signs of degeneration, most of the pollen being abortive. On the other hand, in short styled individuals, the filament, anther, and pollen grains have attained remarkable development; the pollen grians are large and fertile. In short-plant the fertilized flowers readily drop off in every stage of their embryo development. This species has completely lost the self-fertile property, which is characteristic of the non-dimorphic Polygonum genus. Although this specsei typically exhibits the physiological characteristics of the non-dimorphic Polygonum genus. Although this specisei typically exhibits the physiological characteristics of dimorphism in controlled pollination, the short-styled individuals bear no seed in nature, thus misleading taxonomists to idenfity the short-styled plant as male. 2) The morphological feature of the flower organ of P. senticosa obviously indicates definite dimorphism. Physiologically, however, no differentiation towards dimorphism was observed, the species still retaining, both in long and short-individuals, the self-fertile property common to the Polygonum genus. Elaborate examinations revealed that regardless of the modes of pollination, both fertiization and seed setting flourish, no differentiation betwen legitimate and illegitimate unions being recognizable. This sort of physiological property has not been observed in the investigations of other heterostylous plants. It is assumed that this species is differentiated structurally into dimorphism, but not yet physiologically. In nature, however, this plant would have more opportunities to be cross-pollinated, i.e., legitimately combined, than self-pollinated because of the development of two forms of flowers. 3) In terms of heterostylism, the F. esculentum just occupies the intermediate position between P. japonica and P. senticosa structurally, ecologically, and physiologically. Doescription of some of the physiological behavior of the plant will suffice to demonstrate the above facts. While P. japonica has completely lost its self-fertile property, P. senticosa still retains it wolly. In F. esculentum 2-6% of self-fertility is the result in illegitimate combination. There occur occasionally hereditary self fertile individuals among some of the F. or 20 min. irradiation plot, when they reach any stage of the same bacterial population. In addition to this increase of total population in the plots with the more dose of UV light irradiation, it seems that the more dose of UV light irradiation is the more shortened the generation time of Azotobacter. Therefore, it is clear that variation of reproductive rate must be, mere or less, due to the genetic effects induced by UV light irradiation. On the other hand, the lag phase or logarithmic growth phase in nonirradiated culture is shortened prominently, and this must be due to the difference in bacterial number of the original inoculm. The generation time of Azotobacter is shortened by exogeneous treatment of nuclei acid derivatives, and the degree is greater in case of DNA derivatives than RNA dervatives. W.H. Price reported that the rate of ribose nucleic acid to protein in Staphylococcus muscae is proportional to the generation time: that is the faster the cell can form ribose nucleic acid, the more rapid its growth. This explains the shortening of generation time by exogeneous RNA derivatives in this work reasonably. On the other hand, it is well known that the desoxyribose nuclic acid content per cell is constant and independent of the generation time. A.D. Laren and W.N. Takahashi reported that the infectious RNA from TMV is 6 times as sensitive to inactivation by UV as it is in the form of intact virus, and that inactivation of infectious TMV involves onlu a local change on RNA chain. But, the effect of exogeneous DNA in this work suggests that irradiated living cell which cotain DNA bring about some change on DNA moleculs as well as RNA molecules. And if the mutagenic effects of UV take into consideration, it is very reasonable. Therefore, it is clear that the variation of the generation time by UV irradiation is, more or less, due to the genetic effects. Therefore, it seems that the shortness of the average lifewpan of Azotobacter by UV irradiation is resulted not only from the influence of the environmental conditions, but also from the variation of genetic factor of the individual.

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Comparison of Biomass by Forest Fire Type and Recovery at Samcheuk-si, Gangwon-do, Korea (산불 유형별 식생회복정도에 따른 현존생물량 비교)

  • Lim, Seok-Hwa;Kim, Jung-Sup;Shin, Jin-Ho;Bang, Je-Yong;Yang, Keum-Chul
    • Korean Journal of Environment and Ecology
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    • v.26 no.4
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    • pp.528-536
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    • 2012
  • This study has compared the different types of forest fires(unburned, crown fire, ground fire) and the degree of vegetation recovery at Samcheuk-si, Gangwon-do by assessing the biomass and net primary production from July 2007 through July 2010. The research showed that the average biomass of unburned site(Un), crown fire site(C-1), crown fire site(C-3), ground fire site(G-2) were $181.20{\pm}5.39$, $62.04{\pm}4.38$, $131.09{\pm}14.38$, $63.39{\pm}2.72ton{\cdot}ha^{-1}$, respectively. And the research showed that the average net primary production of unburned site(Un), crown fire site(C-1), crown fire site(C-3), ground fire site(G-2) were $4.17{\pm}0.56$, $3.27{\pm}1.56$, $11.51{\pm}0.53$, $2.10{\pm}0.31ton{\cdot}ha^{-1}{\cdot}yr^{-1}$, respectively. Quercus mongolica $DH_{10}$(Diameter at the 10cm tree height) growth rate at each plot was compared to the crown fire site(C-1) in the annual average $1.21{\pm}0.55mm{\cdot}yr^{-1}$ at the speed of the fastest growth follows; showed crown fire site(C-3), ground fire site(G-2), unburned site(Un) appeared in the order. And that showed the growth rate of height was highest in the $15.43{\pm}4.57cm{\cdot}yr^{-1}$ at crown fire site(C-3), then the crown fire site(C-1), and ground fire site(G-2), and lowest in the unburned site(Un).

Effect of Flooding Treatment on the Desalting Efficiency and the Growth of Soiling and Forage Crops in a Sandy Soil of the Iweon Reclaimed Tidal Land in Korea (이원간척지 사질 염류토양의 담수제염처리가 제염효과와 녹비.사료작물의 생육에 미치는 영향)

  • Sohn, Yong-Man;Kim, Hyun-Tea;Jeon, Geon-Yeong;Song, Jae-Do;Lee, Jae-Hwang;Park, Moo-Eon
    • Korean Journal of Soil Science and Fertilizer
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    • v.43 no.1
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    • pp.16-24
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    • 2010
  • Effect of flooding on the desalting efficiency and the growth of sudan grass, barnyard grass, sesbania and corn was studied in a sandy soil of the Iweon reclaimed tidal land. Flooding plots were treated by 400 (one time flooding), 800 (two times flooding), and 1,200 mm(three times flooding) of water, respectively, and then soil salinities of the treated plots were compared with salinity of the control plot (not flooded) for estimation of desalting effect. Desalting ratio of 1,200 mm treatment was 78.3% for depth 0-20 cm, 70.5% for depth 20-40 cm and 60.8% for depth 40-60 cm, and then the soil salinity reached at 3~6 dS $m^{-1}$. Consequently, it was considered that sandy saline soil was satisfactorily desalted for upland crops to be cultivated by 1,200 mm flooding, but insufficiently desalted by 400 mm and 800 mm flooding because of high salinity ranged 5~14 dS $m^{-1}$ even after flooding treatment. In addition, it was estimated that soil salinity should be controled lower than 7.7 dS $m^{-1}$ in order to obtain more than 80%of crop emergence when four crops are simultaneously cultivated by inter- or mixed cropping in a field. Dry matter yields (kg $10a^{-1}$) was 1,068 for sudan grass, 696for barnyard grass, 1,426 for sesbania, and 1,164 for corn by 1,200 mm flooding treatment, but only 46.8~74.3% by 800 mm flooding treatment and 2.9~25.5% by 400 mm flooding treatment. Therefore, it is concluded that the flooding treatment more than 1,200 mm is necessary for satisfactory desalinization in order for the low salt tolerance crops to be cultivated in the sandy reclaimed tidal land.

Effect of nitrogen application and clipping height on the vegetative growth of Korean lawn grass (Zoysia japonica Steud.) and Manilagrass (Zoysia mat rella (L.) MERR.) during September/October (질소시용 및 예초고가 한국 잔디(Zoysia japonica Steud.) 및 금잔디(Zoysia matrella MERR.)의 생육후기 영양생장에 미치는 영향)

  • 심재성;윤익석
    • Asian Journal of Turfgrass Science
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    • v.1 no.1
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    • pp.7-17
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    • 1987
  • The experiment with two levels of nitrogen (0. and 300kg / ha / year) and two levels of clipping height (1.5cm and 4cm) was conducted on the field during the period 3 June to 23 October 1985. Clonal lines of korean lawngrass ( Zoysia japonica Steud.) and manilagrass ( Zoysia matrella ( L.) Merr.)of Daejon origin were established in June, as individual clone in rows 30cm apart with a 40cm spacing between clones, actually 4 clones each plot. The results obtained were as follows : 1. When no nitrogen was applied to korean lawngrass, leaf blade which appeared during the August / early September period remained green for a period of about 10 weeks and even leaves emerged in late September lived for 42 days. However, leaf longevity did not exceed 8 weeks as nitrogen was applied. In contrast the leaf longevity of manilagrass which emerged during the mid - August / early September period was 11 weeks and, under the nitrogen applied, 9 weeks, indicating that the life - saen of individual leaf of manilagrass may be longer than that of korean lawngrass. Meanwhile, clipping height had no effect on the leaf longevity in both grasses. 2. During the July / August period, tiller number, green leaf number and DM weight of korean lawngrass were increased significantly with fertilizer nitrogen, but were not with two levels of clipping height. This trend was reversed after late September : no effect of nitrogen was appeared. Instead, lax clipping increased tiller number, green leaf number and DM weight. Green leaves stimulated by lax clipping resulted in the occurrance of more dead leaves in late October. 3. The increase of tiller number, green leaf number, and DM weight of korean lawngrass due to nitrogen application appeared to be of significance in early September. Unlike korean lawngrass, however, this significant increase was maintained to late October when new green leaves still emerge. Clipping height had little effect on the growth of manilagrass by early September, but since then, lax clipping stimulated leaf appearance, possibly resulting in a remained green color of manilagrass turf. 4. Among the stolons outgrown until early September, the primary stolon was not influenced by nitrogen and clipping treatments to produce only 2 - 3 stolons. However, 1st branch stolon as affected by nitrogen increased significantly, so most of stolons which occurred consisted of 1st branch stolon. 5. Until early September, stolon length obtained at nil nitrogen level was chiefly caused by lengthening the primary stolons. By applying nitrogen the primary stolons of korean lawngrass was longer than 1st branch stolons when severe clipping was involved and in turn, shorter than 1st branch stolons when lax clipping was concerned. In manilagrass, 1st branch stolons were much longer than the primary stolons when turf was clipped severely but in conditions of lax clipping, there was little difference in length between primary and 1st branch stolons. 6. Stolon nodes of both korean lawngrass and manilagrass were positively influenced by nitrogen, but no particular increases by imposing clipping height treatment was marked in manilagrass. Although the stolon of korean lawngrass was grown until late october, the growth stimulated by nitrogen was not so remarkable as to exceed that a by nil N. 7. The thickness of korean lawngrass and manilagrass was greatest in late September, but that of manilagrass did not differ significantly from that in late October. 8. The response of stolon length of korean lawngrass to lax clippings was not so great in late October as to that to severe clippings. On the other hand, the positive effect of lax clippings to stolon length in m anilagrass was confirmed even in late October.

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Effects of Spring Seeding Dates on Growth , Forage Yield and Quality of Early and Late Maturing Oat Cultivars (춘계 파종시기가 조.만생 연맥의 생장 , 사초수량 및 품질에 미치는 영향)

  • 김종림;김동암
    • Journal of The Korean Society of Grassland and Forage Science
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    • v.12 no.2
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    • pp.111-122
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    • 1992
  • This experiment was conducted to determine the effects of spring seeding dates on the growth, yield and quality of early and late maturing spring oat (Auena sativa L.) cultivars on the forage experimental field, College of Agriculture, Seoul National University, Suwon from March to June, 1991. The experiment was arranged as a split plot with three replications. Oat cultivars, Cayuse and Speed oat, were the main plots, and seeding dates consisted of March 15, 22, 29, April 5 and 12 were the subplots. 1. A 7-day delay in seeding represents approximately 3~8 days being early in heading. The heading date of the early maturing cultivar, Speed oat, was 14 days earlier than that of the late maturing cultivar, Cayuse. 2. The concentrations of Crude protein (CP), Acid detergent fiber (ADF), Neutral detergent fiber (NDF) and zn uitro dry matter digestibility (IVDMD) of the late maturing cultivar, Cayuse, harvested May 29 were 19.6, 30.0, 44.9, and 82.7 %, respectively, but those of the early maturing cultivar, Speed oat, were 14.8. 33.3. 52.3, and 71.2 %, respectively. Chemical analyses of oat forage indicated that the contents of crude protein and lVDMD were increased from March 15 to April 12 seeding, while crude fiber was decreased. 3. Theaverage dry matter, IVDDM and CP yields of oats harvested May 29 were 2,960, 2,435 and 572 kg per ha, respectively with the late maturing cultivar. Cayuse. while the early maturing cultivar, Speed oat, recorded 3,255, 2.298 and 475 kg per ha, respectively. No significant dry matter yield differences were found among the different seeding dates of March 15, 22 and 29 for the two oat cultivars. but a significant yield decrease was found from April 5 seeding. No interactions in dry matter yield were observed between oat cultivars and seeding dates. 4. Maximum Leaf area index (LAI) and Leaf area index duration (LAID) were observed with earlier seeding and the LA1 of Cayuse cultivar was twice or three times as much as that of Speed oat cult~var as the growth progresses. 5. As the seeding date was earlier. the Crop growth rate (CGR) of the late maturing cultivar, Cayuse. was increased continuously. but that of the early maturing cultivar, Speed oat, was declined after May 29. This trend was also found on the Net assimilation rate (NAR) of Speed oat cultivar. The present experiment indicates that spring oats can be successfully produced as forages by seeding in March with early maturing cultivars.

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Effect of Seeding Dates of Italian Ryegrass Interseeded into Bermudagrass Sod on the Forage Productivity and Botanical Composition of Italian Ryegrass and Bermudagrass (버뮤다그라스 초지에 보파한 이탈리안 라이그라스의 파종시기가 목초생산성과 식생에 미치는 영향)

  • Park, Nam-Geon;Hwang, Kyung-Jun;Lee, Chong-Eon;Kim, Nam-Young;Park, Hyung-Soo;Ko, Moon-Suk;Lim, Young-Cheol
    • Journal of The Korean Society of Grassland and Forage Science
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    • v.31 no.4
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    • pp.383-388
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    • 2011
  • This study was carried out to determine the effect of seeding dates of Italian ryegrass interseeded into bermudagrass sod on the forage productivity and botanical composition of Italian ryegrass and bermudagrass. Experimental plot was located at 200 m altitude within Subtropical Animal Experiment Station, National Institute of Animal Science in Jeju from 2009 to 2010. Seeding date treatments of Italian ryegrass into bermudagrass sod were arranged in a randomized complete block design replicated three times. Italian ryegrass was drilled in row 20 cm apart after clipping the bermudagrass at a cutting height of 2~3 cm. Seeding time was on 15 September, 30 September and 15 October. The dry matter yield of Italian ryegrass was higher in 15 September seeding treatment, the dry matter yields of 30 September and 15 October seeding treatment were not different. The Italian ryegrass interseeded into bermudagrass sod didn't affect the dry matter yield in the summer harvests of bermudagrass, and the dry matter yield of bermudagrass showed the highest in August. In botanical composition, Italian ryegrass showed to reduced growth in early seedling stage after seeding by competition with bermudagrass, but Italian ryegrass was dominant to June increasing of temperature in spring next year and bermudagrass after Italian ryegrass harvests was dominant during summer season.