DOI QR코드

DOI QR Code

Checklist of the Snake Eels (Ophichthidae: Anguilliformes) from Korea, with a Change of their Korean Names

한국산 바다뱀장어과(개칭)(Ophichthidae: Anguilliformes) 어류의 종 목록 및 국명 개칭

  • Ji, Hwan-Sung (Fisheries Resources Research Center, National Institute of Fisheries Science) ;
  • Kang, Chung-Bae (Department of Exhibition and Education, Marine Biodiversity Institute of Korea) ;
  • Kim, Jin-Koo (Department of Marine Biology, Pukyong National University)
  • 지환성 (국립수산과학원 수산자원연구센터) ;
  • 강충배 (국립해양생물자원관) ;
  • 김진구 (부경대학교 자원생물학과)
  • Received : 2021.11.11
  • Accepted : 2021.12.24
  • Published : 2021.12.31

Abstract

A checklist of snake eels (Ophichthidae) from Korea is presented including information, such as synonyms, distribution, occurrence records and key to species of 7 genera and 14 species. Also, we suggest change of the Korean names for the snake eels (Ophichthidae) "Ba-da-baem-gwa" to "Ba-da-baem-jang-eo-gwa", and it's members, because Korean names of snake eels (Ophichthidae) is similar to those of sea snake (Hydridae).

한국산 뱀장어목 Ophichthidae과 어류 7속 14종의 동종이명, 분포, 출현기록, 종 검색표 등을 정리한 종 목록표를 제시하였다. 또한, Ophichthidae과 어류의 국명이 파충류의 Hydridae과와 유사하여 Ophichthidae과 어류의 국명을 "바다뱀과 어류"에서 "바다뱀장어과 어류"로 개칭하고, 이 과에 속하는 종들의 국명 변경도 함께 제안한다.

Keywords

Acknowledgement

이 논문은 2021년도 국립수산과학원 수산과학연구사업(R2021027) 및 2021년 국립해양생물자원관(2021M01100)의 지원을 받아 수행된 연구입니다.

References

  1. Ahl, J.N. 1789. Specimen ichthyologicum de Muraena et Ophichtho, quod venia exp. fac Med. ups. Praeside Car ol. Pet. Thunberg, etc, Upsalal, pp. 1-14, 2pls.
  2. Allen, G.R. and M. Adrim. 2003. Coral reef fishes of Indonesia. Zool. Stud., 42: 1-72.
  3. Asano, H. 1984. Family Ophichthidae. In: Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino (eds.), The fishes of the Japanese archipelago. Tokai Univ. Press, Tokyo, Japan, pp. 29-32.
  4. Asano, H. 1992. Ophichthidae. In: Masuda, H. (ed.), The fishes of the Japanese archipelago, 3rd ed. Tokai Univ. Press, Tokyo, Japan, pp. 30-32, pl. 33.
  5. Bleeker, P. 1852. Bijdrage tot de kennis der Muraenoiden en Symbrancho den van den Indischen Archipel. Verh. Batav. Genootsch. Kunst. Wet., 25: 1-76.
  6. Bleeker, P. 1853. Diagnostische beschrijvingen van nieuwe of weinig bekende vischsoorten van Batavia. Nat. Tijd. V. Ned. Ind., 4: 451-516.
  7. Castle, P.H.J. and J.E. McCosker. 1999. A new genus and two new species of Myrophine worm-eels, with comments on Muraenichthys and Scolecenchelys (Anguilliformes: Ophichthidae). Rec. Aust. Mus., 51: 113-122. https://doi.org/10.3853/j.0067-1975.51.1999.1300.
  8. Chen, J.T.E. and H.T.C. Weng. 1967. A review of the apodal fishes Taiwan. Tunghai Univ. Taichung, Taiwan, 32: 1-86.
  9. Chen, J.T.F. and M.J. Yu. 1986. A synopsis of the vertebrates of Taiwan, revised and enlarged edition. Commercial Press, Taipei, Taiwan, 1092pp.
  10. Cheng, Q.T. and B.S. Zheng. 1987. Systematic synopsis of Chinese fishes. Science Press, Beijing, pp. 110-115, 775-764.
  11. Chyung, M.K. 1977. The fishes of Korea. Ilji-sa, Seoul, Korea, 727pp.
  12. Eschmeyer, W.N. 2021. Catalog of fishes electronic version. Available at: https://research.calacademy.org/ichthyology/catalog/fishcatmain.asp.
  13. Froese, R. and D. Pauly. 2021. Fishbase. World Wide Web electronic publication. Retrieved from https://www.fishbase.org.
  14. Golani, D. and S.V. Bogorodsky. 2010. The fishes of the Red Sea reappraisal and updated checklist. Zootaxa, 2463: 1-135. https://doi.org/10.11646/zootaxa.2463.1.1.
  15. Gunther, A. 1870. Catalogue of the fishes in the british museum. Catalogue of the Physostomi, containing the families Gymnotidae. in the british museum. Brit. Mus., London, 8: i-xxv+1549pp.
  16. Hatooka, K. 2002. Ophichthidae. In: Nakabo, T.(ed.), Fishes of Japan with pictorial keys to the species. Tokai Univ. Press, Tokyo, Japan, pp. 215-225.
  17. Herre, A.W.C.T. 1923. Check list of Philippine fishes. Res. Rep. U.S. Fish Wild. Serv., 20: 1-977.
  18. Hibino, Y., S. Kimura, K. Hoshino, K. Hatooka and J.E. McCosker. 2012. Validity of Scolecenchelys aoki, with a redescription of Scolecenchelys gymnota (Anguilliformes: Ophichthidae). Ichthyol. Res., 59: 179-188. https://doi.org/10.1007/s10228-011-0269-4.
  19. Hibino, Y., Y. Kai and S. Kimura. 2013. Scolecenchelys fuscogularis(Anguilliformes: Ophichthidae, Myrophinae), a new worm eel from Japan. Ichthyol. Res., 60: 43-47. https://doi.org/10.1007/s10228-012-0309-8.
  20. Hibino, Y. and S. Kimura. 2015. Revision of the Scolecenchelys gymnota species group with descriptions of two new species (Anguilliformes: Ophichthidae: Myrophinae). Ichthyol. Res., 63: 1-22. https://doi.org/10.1007/s10228-015-0485-4.
  21. Hibino, Y., J.E. McCosker and F. Tashiro. 2019a. Four new deepwater Ophichthus(Anguilliformes: Ophichthidae) from Japan with a redescription of Ophichthus pallens (Richardson 1848). Ichthyol. Res., 66: 289-306. https://doi.org/10.1007/s10228-018-00677-3.
  22. Hibino, Y., H.C. Ho and J.E. McCosker. 2019b. A new species Muraenichthys (Anguilliformes: Ophichthidae) from Taiwan, with redescription of Muraenichthys thompsoni Jordan and Richardson, 1908. Zootaxa, 4702: 41-48. https://doi.org/10.11646/zootaxa.4702.1.9.
  23. Hibino, Y. and J.E. McCosker. 2020. Resurrection of Ophichthus zophistius(Actinopterygii: Anguilliformes: Ophichthidae), with a revised diagnosis of O. altipennis. Zootaxa, 4801: 328-342. https://doi.org/10.11646/zootaxa.4801.2.7.
  24. Ho, H.C., D.G. Smith, S.I Wang, K.T. Shao, Y.M. Ju and C.W. Chang. 2010. Specimen catalog of pisces [sic] collection of National Museum of Marine Biology and Aquarium transferred from Tinghai University and Aquarium. (II) Order Anguilliformes. Platax, 7: 13-34.
  25. Ho, H.C. and K.T. Shao. 2011. Annotated checklist and type catalog of fish genera and species described from Taiwan. Zootaxa, 2957: 1-74. https://doi.org/10.11646/zootax.2957.1.1.
  26. Ho, H.C., D.G. Smith, J.E. McCosker, Y. Hibino, K.H. Loh, K.A. Tighe and K.T. Shao. 2015. Annotated checklist of eels (orders Anguilliformes and Saccopharyngiformes) from Taiwan. Zootaxa, 4060: 140-189. https://doi.org/10.11646/zootax.4060.1.16.
  27. Hoshino, K., Y. Hibino, S. Kimura and Y. Machida. 2011. The worm eel, Muraenichthys okamurai Machida and Ohta 1996, a junior synonym of Muraenichthys borealis Machida and Shiogaki 1990. Ichthyol. Res., 58: 184-187. https://doi.org/10.1007/s10228-011-0210-x.
  28. Huang, Z. 2001. Marine species and their distribution in China's seas. Vertebrata. Smithsonian Inst., Florida, U.S.A., 598pp.
  29. Hutchins, J.B. 2001. Checklist of the fishes of Western Australia. Rec. West. Aust. Mus. Suppl., 63: 9-50. https://doi.org/10.18195/issn.0313-122x.63.2001.009-050
  30. Jeong, C.H. 1999. A review of taxonomic studies and common names of rajid fishes(Elasmobranchii, Rajidae) from Korea. Korean J. Ichthyol., 11: 198-210.
  31. Ji, H.S. 2014. Phylogenetic study of Anguilliformes species based on leptocephali from the adjacent waters of Korea. Doctoral Thesis. Pukyong National University, Busan, Korea, 204pp.
  32. Ji, H.S. and J.K. Kim. 2010. Molecular and morphological identification of ophichthid leptocephali from the south sea of Korea. Korean J. Ichthyol., 22: 279-284.
  33. Ji, H.S. and J.K. Kim. 2011a. Taxonomic review of the snake-eels family Ophichthidae (Anguilliformes) from Korea. Korean J. Ichthyol., 23: 46-60.
  34. Ji, H.S. and J.K. Kim. 2011b. A new species of snake eel, Pisodonophis sangjuensis(Anguilliformes: Ophichthidae) from Korea. Zootaxa, 2758: 57-68. https://doi.org/10.11646/zootaxa.2758.1.4.
  35. Ji, H.S., S.J. Lee and J.K. Kim. 2011. Molecular identification, ontogeny and evolutionary note of Echelus uropterus leptocephali. Korean J. Ichthyol., 23: 217-224.
  36. Ji, H.S., H.W. Lee, B.K. Hong and J.K. Kim. 2012. Four newly reported ophichthid leptocephali species revealed by mitochondrial 12S rDNA, with implications of their occurrence in Korea. Anim. Cells Syst., 16: 415-424. https://doi.org/10.1080/19768354.2011.652168.
  37. Ji, H.S. and J.K. Kim. 2012. Molecular and morphological identification of Muraenichthys gymnopterus(Ophichthidae: Anguilliformes) leptocephalus collected from Jeju Island, Korea. Kor. J. Fish. Aquat. Sci., 45: 507-512. https://doi.org/10.5657/KFAS.2012.0507.
  38. Ji, H.S., J.H. Choi, K.H. Choi, D.W. Lee and J.K. Kim. 2013a. First morphological description of Ophichthus asakusae (Ophichthidae: Anguilliformes) leptocephali collected from the waters south of Jeju Island. Kor. J. Fish. Aquat. Sci., 46: 594-597. https://doi.org/10.5657/KFAS.2013.0594.
  39. Ji, H.S., M.J. Kim and J.K. Kim. 2013b. First description of the adult Scolecenchelys aoki(Ophichthidae, Anguilliformes) collected from the South Sea of Korea, with comments on the synonym Scolecenchelys borealis. Kor. J. Fish. Aquat. Sci., 46: 589-593. https://doi.org/10.5657/KFAS.2013.0589.
  40. Ji, H.S., J.H. Choi, K.H. Choi, S.C. Yoon, D.W. Lee and J.K. Kim. 2014. First morphological description and the distribution of Ophisurus macrorhynchos (Anguilliformes: Ophichthidae) leptocephalus collected from southeastern waters of Jeju Island. Kor. J. Fish. Aquat. Sci., 47: 888-894. https://doi.org/10.5657/KFAS.2014.0888.
  41. Ji, H.S., J.K. Kim, S.J. Lee, S. Kimura and Y. Hibino. 2015. New record of Scolecenchelys fuscogularis(Anguilliformes: Ophichthidae) leptocephali from Korea, as revealed by morphological and molecular analyses. Fish. Aquat. Sci., 18: 109-113. https://doi.org/10.5657/FAS.2015.0109.
  42. Jordan, D.S. and J.O. Snyder. 1901. A review of the apodal fishes or eels of Japan, with descriptions of nineteen new species. Proc. U.S. Natl. Mus., 23: 837-890. https://doi.org/10.5479/si.00963801.23-1239.837
  43. Jordan, D.S. and R.E. Richardson. 1908. Fishes from islands of the Philippine Archipelago. Bull. Bure. Fish., 27: 233-287.
  44. Jordan, D.S. and R.E. Richardson. 1909. A catalogue of the fishes of the island of Formosa, or Taiwan, based on the collections of Dr. Hans Sauter. Mem. Carn. Mus., 4: 159-204, pls. 63-74.
  45. Jordan, D.S. and R.E. Richardson. 1910. Check-list of the species of fishes known from the Philippine archipelago. Bureau of Sci., Manila, Philippines, pp. 3-78.
  46. Kaup, J.J. 1856. Uebersicht der Aale. Archiv fur Naturgeschichte, 22: 41-77. https://doi.org/10.5962/bhl.part.11240
  47. Kang, Y.S., I.B. Yoon. 1975. Illustrated Encyclopedia of fauna and flora of Korea. Vol. 17. Amphibia and Reptilia. Sam-Hwa Press, Seoul, Korea, 190pp.
  48. Kang, C.B., J.S. Kim, J.K. Kim, H.S. Ji, M.G. Yoon and J.W. Park. 2019. A new record of the family Ophichthidae (Pisces), Ophichthus erabo (Jordan and Snyder 1901), from Korea. Korean J. Fish. Aquat. Sci., 52: 418-423. https://doi.org/10.5657/KFAS.2019.0418.
  49. Kim, B.G., C.H. Jeong and K.N. Han. 2008. New record of a worm eel Muraenichthys gymnopterus(Anguilliformes: Ophichthidae) from Korea. Korean J. Ichthyol., 20: 318-323.
  50. Kim, B.J., J.H. Choi, D.S. Chang, H.K. Cha and J.H. Park. 2009. New record of the snake eel Ophichthus asakusae (Ophichthidae: Anguilliformes) from Korea. Fish. Aqua. Sci., 12: 236-239. https://doi.org/10.5657/fas.2009.12.3.236.
  51. Kim, I.H., J.J. Park, W.J. Choi, K.S. Koo and D.S. Park. 2017. Analysis of the correct and incorrect reports on the observation and capture of sea snake in the coasts of South Korea. Kor. J. Environ. Ecol., 31: 455-460. https://doi.org/10.13047/KJEE.2017.31.5.455.
  52. Kim, I.S., Y. Choi, C.L. Lee, Y.J. Lee, B.J. Kim and J.H. Kim. 2005. Illustrated book of Korean fishes. Kyohak Publishing Co. Ltd., Seoul, Korea, 615pp.
  53. Kim, J.K., J.H. Ryu, H.J. Kwun, H.S. Ji, J.H. Park, S.H. Myoung, Y.S. Song, S.J. Lee, H.J. Yu, S.E. Bae, S.H. Jang and W.J. Lee. 2019. Distribution map of Sea fishes in Korean peninsula. Ministry of Oceans and Fisheries, Korea Institute of Marine Science and Technology Promotion and Pukyong National University, 541pp.
  54. Lacepede, B.G.E. 1800. Histoire naturelle des poissons. Histoire naturelle des poissons, 2: i-lxiv+632pp, Pls. 1-20.
  55. Lee, C.L. 2009. First record of a longfin snake-eel, Pisodonophis cancrivorus(Anguilliformes: Ophichthidae) in Korea. Korean J. Ichthyol., 21: 307-310.
  56. Lee, C.L. and H. Asano. 1997. A new ophichthid eel, Ophichthus rotundus(Ophichthidae, Anguilliformes) from Korea. Kor. J. Biol. Sci., 1: 549-552.
  57. Linnaeus, C. 1758. Systema Naturae, Ed. X. (Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.) Holmiae. Systema Naturae, Ed. X. v. 1: i-ii+824pp.
  58. Machida, Y. and M. Shiogaki. 1990. A new snake eel, Muraenichthys borealis, from Aomori, northern Japan. Jpn. J. Ichthyol., 37: 1-5.
  59. Machida, Y. and S. Ohta. 1993. New record for Neenchelys daedalus (Ophichthidae) from Japan. Jpn. J. Ichthyol., 39: 391-394. https://doi.org/10.1007/BF02905140
  60. Machida, Y. and S. Ohta. 1996. Description of a new worm eel, Muraenichthys okamurai, from western Honshu, Japan (Ophichthidae: Myrophinae). Mem. Fac. Sci. Kochi Univ., 16/17: 77-81.
  61. Macleay, W. 1881. A descriptive catalogue of Australian fishes. Part IV. Proc. Linn. Soc. NSW., 6: 202-387.
  62. Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino. 1984. The fishes of the Japanese Archipelago. Tokyo, Japan, Text: i-xxii+437pp, Pls. 1-370.
  63. McCosker, J.E. 1970. A review of the eel genera Leptenchelys and Muraenichthys, with the description of a new genus, Schismorhynchus, and a new species, Muraenichthys chilensis. Pacific Sci., 24: 506-516.
  64. McCosker, J.E. 1977. The osteology, classification, and relationships of the eel family Ophichthidae. Proc. Calif. Acad. Sci. Ser. 4, 41: 1-123.
  65. McCosker, J.E. 2010. Deepwater Indo-Pacific species of the snakeeel genus Ophichthus (Anguilliformes: Ophichthidae), with the description of nine new species. Zootaxa, 2505: 1-139. https://doi.org/10.11646/zootaxa.2505.1.1.
  66. McCosker, J.E. and P.H.J. Castle. 1986. Ophichthidae. In: Smith, M.M. and P.C. Heemstra (eds.), Smiths' Sea fishes. Springer-Verlag, Berlin, Germany, pp. 176-186.
  67. McCosker, J.E. and Y.Y. Chen. 2000. A new species of deepwater snake-eel, Ophichthus aphotistos, with comments on Neenchelys retropinna (Anguilliformes: Ophichthidae) from Taiwan. Ichthyol. Res., 47: 353-357. https://doi.org/10.1007/BF02674262.
  68. McCosker, J.E. and J.E. Randall. 2001. Revision of snake-eel genus Brachysomophis (Anguilliformes: Ophichthidae), with description of two new species and comments on the species of Mystriophis. Indo-Pacific Fish., 33: 1-32.
  69. McCosker, J.E., G.R. Allen, D.F. Hoese, J.E. Gates and D.J. Bray. 2006. Family Ophichthidae. In: Hoese, D.F., D.J. Bray, J.R. Paxton and G.R. Allen (eds.), Zoological Catalogue of Australia. Vol. 35. Pisces. Australian Government Publishing Service, Canberra, Australia, pp. 264-277.
  70. McCosker, J.E. and H.C. Ho. 2014. Redescription of the sharp-fanged snake eel Brachysomophis longipinnis (Anguilliformes: Ophichthidae). Platax, 11: 95-101.
  71. McCosker, J.E. and H.C. Ho. 2015. New species of the snake eels Echelus and Ophichthus (Anguilliformes: Ophichthidae) from Taiwan. Zootaxa, 4060: 71-85. https://doi.org/10.11646/zootaxa.4060.1.11.
  72. McCosker, J.E., S.V. Bogorodsky, A.O. Mal and T.J. Alpermann. 2020. Description of a new snake eel Ophichthus olivaceus (Teleostei: Anguilliformes, Ophichthidae) from the Red Sea. Zootaxa, 4750: 31-48. https://doi.org/10.11646/zootaxa.4750.1.2.
  73. Mohapatra, A., D. Ray, S.R. Mohanaty and S.S. Mishra. 2018. Ophichthus johnmccoskeri sp. nov.(Anguilliformes: Ophichthidae): a new snake eel from Indian waters, Bay of Bengal. Zootaxa, 4462: 251-256. https://doi.org/10.11646/zootaxa.4462.2.7
  74. Mohapatra, A., D. Ray, S.R. Mohanaty and S.S. Mishra. 2020. First report of a snake eel, Ophichthus sangjuensis (Ji and Kim, 2011), (Anguilliformes: Ophichthidae), from Indian Ocean. Thalassas: An Int. J. Mar. Sci., 36, 553-555. https://doi.org/10.1007/s41208-020-00240-w.
  75. Mohapatra, A., S.R. Mohanty, D. Ray, S.S. Mishra and J.K. Seth. 2021. A new species of the genus Cirrhimuraena (Anguilliformes: Ophichthidae) from the Bay of Bengal, India. J. Fish. Biol., 98: 1363-1370. https://doi.org/10.1111/jfb.14671.
  76. Mok, H.K. 1993. Ophichthidae In: Shen, S.C.(ed.), Fishes of Taiwan, Depart. Zool., Natl. Taiwan Univ., Taipei, pp. 110-114.
  77. Mori, T. 1952. Check list of the fishes of Korea. Mem. Hyogo Univ. Agric., 1: 1-228.
  78. Myers, R.F. 1999. Micronesian reef fishes. A comprehensive guide to the coral reef fishes of Micronesia, 3rd ed. Coral Graphics, Barrigada, Guam, i-vi+330pp, 192pls.
  79. Nelson, J.S., T.C. Grande and V.H. Wilson. 2016. Fishes of the world, 5th ed, John Wiley & Sons Inc., New Jersey, U.S.A., 707pp.
  80. Ogilby, J.D. 1897. Some new genera and species of fishes. Proc. Linn. Soc. NSW., 22: 245-251.
  81. Rafinesque, C.S. 1810. Caratteri di alcuni nuovi generi e nuove specie di animali e piante della Sicilia, con varie osservazioni sopra i medisimi. Part 1 involves fishes, pp. 3-69.
  82. Randall, J.E. and K.K.P. Lim. 2000. A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl., 8: 569-667.
  83. Richardson, J. 1848. Ichthyology. In: Richardson, J. and J.E. Gray (eds.), The zoology of the voyage of H.M.S. "Erebus" and "Terror" under the command of Capt Sir J.C. Ross during 1839-43. London, 2: 1-139, 62pls.
  84. Shao, K.T., H.C. Ho, P.L. Lin, P.F. Lee, M.Y. Lee, C.Y. Tsai, Y.C. Liao and Y.C. Lin. 2008. A checklist of the fishes of southern Taiwan, Northern South China Sea. Raffles Bull. Zool. Suppl., 19: 233-271.
  85. Shen, S.C. 1984. Synopsis of fishes of Taiwan. Southern Materials Center, Taipei, Taiwan, 533pp.
  86. Shen, S.C., S.C. Lee, K.T. Shao, H.K. Mok, C.T. Chen and C.H. Chen. 1993. Fishes of Taiwan. Department of Zoology, Natl. Taiwan Univ., Taipei, Taiwan, 960pp.
  87. Smith, D.G. and J.E. McCosker. 1999. Ophichthidae. In: Carpenter, K.E. and V.H. Niem (eds.), FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 3. Batoid fishes, chimaeras and bony fishes. Part 1 (Elopidae to Linophrynidae), FAO, Rome, Italy, pp. 1662-1669.
  88. Soniyama, T., K. Ogimoto, S. Hori, Y. Uchida and M. Kawano. 2020. An annotated checklist of marine fishes of the Sea of Japan off Yamaguchi Prefecture, Japan, with 74 new records. Bull. Kagoshima Univ. Mus., 11: 1-152.
  89. Sumida, S. and Y. Machida. 2000. Revision of the two sympatric snake-eel species of the genus Ophichthus (Ophichthidae, Anguilliformes) from Tosa Bay, southern Japan, with comments on O. tsuchidae. Bull. Mar. Sci. Fish., Kochi Univ., 20: 51-69.
  90. Tang, W.Q. and C.G. Zhang. 2004. A taxonomic study on snake eel family Ophichthidae in China with the review of Ophichthidae (Pisces, Anguilliformes). J. Shanghai Fish. Univ., 13: 16- 22 (in Chinese). https://doi.org/10.3969/j.issn.1004-7271.2004.01.004
  91. Temminck, C.J. and H. Schlegel. 1846. Pisces. In: Siebold, P.F. de (ed.), Fauna Japonica, sive descriptio animalium, quae in itinere per Japoniam suscepto annis 1823-1830 collegit, notis, observationibus et adumbrationibus illustravit Ph.Fr. de Siebold, Lugduni Batavorum, Parts 10-14: 173-269.
  92. Vo, Q.V., Y. Hibino and H.C. Ho. 2019. A new species of the snake eel genus Ophichthus, with additional records from Viet Nam (Anguilliformes: Ophichthidae). Zool. Stud., 58: 43: 1-14. https://doi.org/10.26107/RBZ-2021-0006.
  93. Zhang, C.G., W.Q. Tang, D. Liu, Z.L. Zhang and S.Y. Zhang. 2010. Fauna Sinica. Osteichthyes. Anguilliformes, Notacanthiformes. Science Press, Beijing, China, i-x+1-453pp., Pls. 1.1, 1.2, 2-3.