Acknowledgement
This study was supported financially by the Agricultural Animal Breeding Project of Shandong Province (No. 2020LZGC012), Funds of Shandong "Double Tops" Program (No. SYL2017YSTD12), Shandong Modern Pig Technology & Industry System Project (No. SDAIT-08-02), Shandong Provincial Natural Science Foundation (No. ZR2018BC046, ZR2019MC053).
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